Coptoclavidae

Ditomoptera dubia

Taxonavigation

Taxonavigation: Dytiscoidea

Superregnum: Eukaryota Cladus: Unikonta Cladus: Opisthokonta Cladus: Holozoa Regnum: Animalia Subregnum: Eumetazoa Cladus: Bilateria Cladus: Nephrozoa Cladus: Protostomia Cladus: Ecdysozoa Cladus: Panarthropoda Phylum: Arthropoda Cladus: Pancrustacea Cladus: Allotriocarida Subphylum: Hexapoda Classis: Insecta Cladus: Dicondylia Subclassis: Pterygota Infraclassis: Neoptera Cladus: Eumetabola Cladus: Holometabola Superordo: Coleopterida Ordo: Coleoptera Subordo: Adephaga Superfamilia: Dytiscoidea

Familia: †Coptoclavidae
Subfamiliae: †Charonoscaphinae – †Coptoclavinae – †Coptoclaviscinae – †Hispanoclavinae – †Timarchopsinae
Genera incertae subfamiliae: †Amblycephalonius – †Carabopteron

Name

Coptoclavidae Ponomarenko, 1961.

Type genus: Coptoclava Ping, 1928.

References

Primary references

• Ponomarenko, A.G. 1961. О систематическом положении Coptoclava longipoda Ping (Insecta, Coleoptera) [About the systematic position of Coptoclava longipoda Ping (Insecta, Coleoptera)]. Paleontologicheskii Zhurnal 1961(3): 67–72. (in Russian). PDF. Reference page. 

Additional references

• Beutel, R.G., Wang, B., Tan, J.-J., Ge, S.-Q., Ren, D. & Yang, X-K. 2012. On the phylogeny and evolution of Mesozoic and extant lineages of Adephaga (Coleoptera, Insecta). Cladistics 29(2): 147–165. DOI: 10.1111/j.1096-0031.2012.00420.x . Reference page. 
• Ponomarenko, A.G., Prokin, A.A. & Bashkuev, A.S. 2015. Coptoclavid beetles (Insecta: Coleoptera: Adephaga) from the Triassic of Lower Franconia, Germany. Paleontological Journal 49(12): 1334–1345. DOI: 10.1134/S0031030115120096 . PDF. Reference page. 
• Prokin, A.A., Makarov, K.V., Ponomarenko, A.G. & Bashkuev, A.S. 2013. New beetle larvae (Coleoptera: Coptoclavidae, Caraboidea, Polyphaga) from the Upper Triassic of Germany. Russian Entomological Journal 22(4): 259–274. PDF. Reference page. 
• Soriano, C., Ponomarenko, A.G. & Delclòs, X. 2007. Coptoclavid beetles (Coleoptera: Adephaga) from the lower Cretaceous of Spain: a new feeding strategy in beetles. Palaeontology 50(2): 525–536. DOI: 10.1111/j.1475-4983.2007.00642.x . Reference page. 
• Wang, B., Ponomarenko, A.G. & Zhang, H.-Ch. 2009. [A new coptoclavid larva (Coleoptera: Adephaga: Dytiscoidea) from the Middle Jurassic of China, and its phylogenetic implication]. Paleontologicheskii Zhurnal 2009(6): 49–57. [in Russian, English translation in Paleontological Journal 43(6): 652–659. DOI: 10.1134/S0031030109060082 .] Reference page. 
• Wang, B., Ponomarenko, A.G. & Zhang, H. 2010. Middle Jurassic Coptoclavidae (Insecta: Coleoptera: Dytiscoidea) from China: a good example of mosaic evolution. Acta Geologica Sinica (English Edition) 84(4): 680–687. DOI: 10.1111/j.1755-6724.2010.00272.x . Reference page. 

Links

• Coptoclavidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD
• Maddison, David R. 1995. Coptoclavidae. Version 01 January 1995 (under construction) in The Tree of Life Web Project