Mikhail Mikhailovich Omelko

Mikhail Mikhailovich Omelko, Russian entomologist.

• Gornotaezhnaya Station FEB RAS, Gornotaezhnoe Vil., Ussuriyski Dist., Primorski krai 692533, Russia. E-mail: omelkom@gmail.com
• Far Eastern Federal University, Sukhanova 8, Vladivostok 690950, Russia
• E-mail: mmomelko@mail.ru

Taxon names authored

(List may be incomplete)

• 146 taxon names authored by Mikhail Mikhailovich Omelko

Publications

(List may be incomplete)

1984

• Omelko, M.M. 1984. Description of three new species of Gelechiidae from South Primorye. In: Kononenko VS, Kupyanskaya AN, Ler PA (Eds) Fauna i ecologia nasekomykh yuga Dal'nego [Fauna and Ecology of far East Insects]. 112(215): 28–34. [In Russian]. Reference page. 

1988

• Omelko, M.M. 1988. New genera and species of Gelechiid moths of the tribe Gelechiini (Lepidoptera, Gelechiidae) from South Primorye Entomologicheskoe Obozrenie 67(1): 142–159. Reference page. 

1993

• Omelko, M.M. 1993. Gelechiid moths of the genus Thiotricha Meyr. (Lepidoptera, Gelechiidae) of the Primorye Territory. Biological Investigations Gornot. Stn. Russian Academy Sciences 201–215, 241–251 [In Russian]. Reference page. 

2003

• Ponomarenko, M.G. & Omelko, M.M. 2003. Review of the genus Acanthophila Heinemann, 1870 (Lepidoptera, Gelechiidae). Far Eastern Entomologist 127: 1–24. PDF Reference page. 

2012

• Marusik, Yu.M., Zhang, F. & Omelko, M.M. 2012: A new genus and species of ctenid spiders (Aranei: Ctenidae) from South China. Arthropoda selecta 21(1): 61–66. Full article (PDF). Reference page. 
• Omelko, M.M. & Marusik, Yu.M. 2012. A review of the Haplodrassus montanus-group (Aranei: Gnaphosidae) in the East Palaearctic and preliminary grouping of the genus. Arthropoda selecta 21(4): 339–348. Full article (PDF). Reference page. 
• Storozhenko, S.Yu. & Omelko, M.M. 2012. Review of the genus Pseudoxistrella Liang, 1991 (Orthoptera: Tetrigidae, Metrodorinae). Zootaxa 3573: 59–65. Preview Reference page. 

2013

• Fomichev, A.A., Marusik, Yu.M. & Omelko, M.M. 2013. A new species of Gnaphosa Latreille, 1804 (Aranei: Gnaphosidae) from western Mongolia. Arthropoda Selecta 22(2): 153-156. DOI: 10.15298/arthsel.22.2.04. . Reference page. 
• Kunt, K.B., Marusik, Yu.M. & Omelko, M.M. 2013. A review of the genus Hygrocrates with a description of a new species from Turkey (Araneae: Dysderidae). Acta Zoologica Bulgarica 65: 273–277. Reference page. 
• Marusik, Y.M., Nadolny, A.A. & Omelko, M.M. 2013: A survey of East Palaearctic Lycosidae (Araneae). 10. Three new Pardosa species from the mountains of Central Asia. Zootaxa 3722(2): 204–218. DOI: 10.11646/zootaxa.3722.2.5. Reference page. 
• Marusik, Yu.M., Omelko, M.M. & Benjamin, S.P. 2013. The first description of adult female of Borboropactus asper (O. P.-Cambridge, 1884) from Sri Lanka (Araneae: Thomisidae). Zootaxa 3737(2): 197–200. DOI: 10.11646/zootaxa.3737.2.8. Reference page. 
• Marusik, Y.M., Omelko, M.M. & Ryabukhin, A.S. 2013. New data on spiders (Aranei) from Eastern Koryakia, Kamchatka Peninsula. Arthropoda selecta 22(4): 363–377.
• Omelko, M.M. & Marusik, Y.M. 2013. A survey of East Palaearctic Lycosidae (Araneae). 9. A review of Sibirocosa with a description of three new species. Zootaxa 3666(3): 319–330. DOI: 10.11646/zootaxa.3666.3.2. . Reference page. 
• Omelko, M.M., Marusik, Y.M. & Koponen, S. 2013. A new species of Alopecosa Simon (Araneae, Lycosidae) from Sakhalin Island. Acta zootaxonomica Sinica 38(2): 267–271. Abstract. Reference page. 

2014

• Kovblyuk, M.M., Kastrygina, Z.A. & Omelko, M.M. 2012: New Lathys Simon, 1884 species from Crimea (Aranei: Dictynidae). Arthropoda Selecta 23(2): 195–198. Full article (PDF). Reference page. 
• Kronestedt, T., Marusik, Yu.M. & Omelko, M.M. 2014. Studies on species of Holarctic Pardosa groups (Araneae, Lycosidae). VIII. The Palearctic species of the Pardosa nigra group. Zootaxa 3894(1): 33–60. DOI: 10.11646/zootaxa.3894.1.5 Reference page. 
• Marusik, Yu.M., Fomichev, A.A. & Omelko, M.M. 2014: A survey of East Palaearctic Gnaphosidae (Araneae). 2. Two new Gnaphosa Latreille, 1804 species from Western Mongolia. ZooKeys 426: 1–9. DOI: 10.3897/zookeys.426.7898 Reference page. 
• Marusik, Yu.M., Fomichev, A.A. & Omelko, M.M. 2014a. A survey of East Palaearctic Gnaphosidae (Araneae). 1. On the Berlandina Dalmas, 1922 (Gnaphosinae) from Mongolia and adjacent regions. Zootaxa 3827(2): 187–213. DOI: 10.11646/zootaxa.3827.2.4. . Reference page. 
• Marusik, Y.M., Nadimi, A., Omelko, M.M. & Koponen, S. 2014. First data about cave spiders (Arachnida: Araneae) from Iran. Zoology in the Middle East 60(3): 255–266. DOI: 10.1080/09397140.2014.943465 Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2014. A survey of East Palaearctic Gnaphosidae (Araneae). 3. On new and poorly known Gnaphosa Latreille, 1804. Zootaxa 3894(1): 10–32. DOI: 10.11646/zootaxa.3894.1.4 Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2014. Reconsideration of Xysticus species described by Ehrenfried Schenkel from Mongolia and China in 1963 (Araneae: Thomisidae). Zootaxa 3861(3): 275–289. DOI: 10.11646/zootaxa.3861.3.5 Reference page. 
• Šestáková, A., Marusik, Yu.M. & Omelko, M.M. 2014. A revision of the Holarctic genus Larinioides Caporiacco, 1934 (Araneae: Araneidae). Zootaxa 3894(1): 61–82. DOI: 10.11646/zootaxa.3894.1.6 Reference page. 
• Yu, H., Zhang, F. & Omelko, M.M. 2014. A new species of Diphya Nicolet 1849 and the first description of the female of D. tanasevitchi from northern China (Araneae: Tetragnathidae: Metinae). Acta Arachnologica 63(1): 31–35. DOI: 10.2476/asjaa.63.31 Reference page. 

2015

• Marusik, Y.M., Azarkina, G.N. & Omelko, M.M. 2015. Redescription and a new synonym of the spider Pachygnatha amurensis Strand, 1907 (Araneae, Tetragnathidae, Tetragnathinae). Zootaxa 4007(1): 126–130. DOI: 10.11646/zootaxa.4007.1.10. Preview (PDF) Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2015. Redescription of "Amaurobius" rufipes Taczanowski, 1874 (Araneae: Corinnidae) with notes of Amaurobius species described from French Guyana. Zootaxa 3974(1): 148–150. DOI: 10.11646/zootaxa.3974.1.14 Preview (PDF). Reference page. 
• Marusik, Yu.M., Omelko, M.M. & Koponen, S. 2015. A survey of East Palaearctic Lycosidae (Araneae). 11. Two new genera from the Acantholycosa complex. Zootaxa 3985(2): 252–264. DOI: 10.11646/zootaxa.3985.2.4. Preview (PDF). Reference page. 
• Marusik, Yu.M., Omelko, M.M. & Koponen, S. 2015. A redescription of the ant mimicking spider Myrmecium gounellei (Araneae: Corinnidae, Castianeirinae), with notes on the genus. Zootaxa 3985(2): 296–300. DOI: 10.11646/zootaxa.3985.2.9 Preview (PDF). Reference page. 
• Marusik, Yu.M., Omelko, M.M. & Koponen, S. 2015. Redescription of “Nesticus” citrinus (Taczanowski, 1874) (Araneae, Araneoidea) from French Guyana. Zookeys 537: 97–101. DOI: 10.3897/zookeys.537.6818 Full article. Reference page. 
• Marusik, Y.M., Zonstein, S.L. & Omelko, M.M. 2015. Redescription of Phrurolithus flavipes (Araneae: Phrurolithidae), with the first description of the male. Zootaxa 4020(1): 197–200. DOI: 10.11646/zootaxa.4020.1.10 Preview (PDF). Reference page. 

2016

• Fomichev, A.A., Marusik, Yu.M. & Koponen, S. 2016. On the synonymy of two Acantholycosa species (Araneae, Lycosidae) from the Altai. ZooKeys 559: 151–156. DOI: 10.3897/zookeys.559.7048. Reference page. 
• Marusik, Y.M. & Omelko, M.M. 2016. Redescription of Draposa subhadrae (Patel & Reddy, 1993) (Araneae: Lycosidae, Pardosinae). Zootaxa 4107(1): 89–92. DOI: 10.11646/zootaxa.4107.1.6. Reference page. 
• Marusik, Y.M. & Omelko, M.M. 2016. Redescription of two Ctenidae (Araneae) from Taiwan. Zootaxa 4173(2): 155–162. DOI: 10.11646/zootaxa.4173.2.6. Reference page. 
• Marusik, Y.M., Omelko, M.M. & Koponen, S. 2016. Two new Bifurcia species (Aranei: Linyphiidae: Micronetinae) from Far East Russia. Arthropoda Selecta 25(1): 77-83. DOI: 10.15298/arthsel.25.1.08. . Reference page. 
• Nadolny, A.A., Omelko, M.M., Marusik, Yu.M. & Blagoev, G. 2016. A new species of spider belonging to the Pardosa lugubris-group (Araneae: Lycosidae) from Far East Asia. Zootaxa 4072(2): 263–281. DOI: 10.11646/zootaxa.4072.2.8. Reference page. 
• Omelko, M.M., Komisarenko, A.A. & Marusik, Yu.M. 2016. A new species of Acantholycosa Dahl, 1908 (Araneae: Lycosidae) from the Russian Far East. Zootaxa 4072(5): 596–600. DOI: 10.11646/zootaxa.4072.5.9. Reference page. 
• Zonstein, S.L., Marusik, Yu.M. & Omelko, M.M. 2016. Redescription of the type species of Diaphorocellus Simon, 1893 (Araneae, Palpimanidae, Chediminae). African Invertebrates 57(2): 93–103. DOI: 10.3897/AfrInvertebr.57.9988. Reference page. 

2017

• Marusik, Yu.M. & Omelko, M.M. 2017. A new species of Acantholycosa baltoroi-group (Araneae: Lycosidae, Pardosinae) from the Russian Far East. Zootaxa 4232(4): 597–600. DOI: 10.11646/zootaxa.4232.4.12 Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2017a. Redescription of Micaria beaufortia (Araneae, Gnaphosidae), with notes on Afrotropical Micaria. Zootaxa 4294(4): 462–470. DOI: 10.11646/zootaxa.4294.4.5. Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2017. A new species of Tekellina (Araneae, Araneoidea) from the Russian Far East. Entomologica Fennica 28(3): 164-168. DOI: 10.33338/ef.84684 Full article (PDF). Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2017. A new species of Acantholycosa baltoroi-group (Araneae: Lycosidae, Pardosinae) from the Russian Far East. Zootaxa 4232(4): 597–600. DOI: 10.11646/zootaxa.4232.4.12 Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2017d. A new genus of tetragnathid spiders from Papua New Guinea (Aranei, Tetragnathidae). Vestnik Zoologii 51(3): 203-214. DOI: 10.1515/vzoo-2017-0027. . Reference page. 
• Marusik, Yu.M., Omelko, M.M. & Ponomarev, A.V. 2017. A survey of the Holarctic genus Arctella Holm, 1945 (Araneae: Dictynidae, Tricholathysinae), with the description of Tricholathys ovtchinnikovi sp. n. Oriental Insects 51(3): 246-261. DOI: 10.1080/00305316.2017.1279086. . Reference page. 

2018

• Marusik, Yu.M. & Omelko, M.M. 2018. New data on Synaphosus (Araneae: Gnaphosidae) from Southeast Asia. Zootaxa 4374(2): 235–248. DOI: 10.11646/zootaxa.4374.2.4 Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2018. A survey of the Porrhoclubiona Lohmander, 1944 from Central Asia (Araneae, Clubiondae). ZooKeys 802: 19–38. DOI: 10.3897/zookeys.802.30236. Reference page. 
• Marusik, Yu.M., Omelko, M.M. & Koponen, S. 2018. Redescription of the Himalayan Trachelas costatus (Araneae: Trachelidae). Zootaxa 2018(4433): 2–390. DOI: 10.11646/zootaxa.2018.4433.392 Reference page. 
• Omelko, M.M. & Omelko, N.V. 2018. New species of the genus Photodotis Meyrick (Lepidoptera, Gelechiidae) in the fauna of Laos. Entomological Review 98(2): 208–212. (Original Russian Text published in Zoologicheskii Zhurnal, 2018, 97(1): 17–22.) DOI: 10.1134/S0013873818020112 . Reference page. 
• Omelko, M.M. & Omelko, N.V. 2018a. A New Genus and Two New Species of Gelechiid Moths of the Subfamily Anacampsinae (Lepidoptera, Gelechiidae) from Laos. Entomological Review 98(9)): 1323–1328. (Original Russian Text published in Zoologicheskii Zhurnal, 2018, 97(11): 1394–1399. DOI: 10.1134/S0013873818090105 . Reference page. 
• Omelko, M.M. & Omelko, N.V. 2018b. NEW FINDS OF THE GELECHIID OF SUBFAMILY ANOMOLOGINAE (LEPIDOPTERA, GELECHIIDAE) IN THE PRIMORSKII KRAI. 4. THE GENUS MONOCHROA HEINEMANN, 1870 (GROUP CONSPERSELLA AND PALUSTRELLA). Amurian zoological journal X(2): 111-122. (in Russian). Reference page. 
• Zonstein, S.L., Marusik, Yu.M. & Omelko, M.M. 2018. Redescription of the monotypic genus Scelidomachus Pocock, 1899 (Aranei: Palpimanidae) and its type species. Arthropoda Selecta 27(1): 53–56. Full article (PDF). Reference page. 

2019

• Marusik, Yu.M., Fomichev, A.A. & Omelko, M.M. 2019. New data on Parasyrisca (Araneae: Gnaphosidae) from Mongolia. Zootaxa 4688(2): 199–212. DOI: 10.11646/zootaxa.4688.2.2 . Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2019. Redescription of gnaphosids (Aranei: Gnaphosidae) described by O. Pickard-Cambridge from the material of the Second Yarkand Mission. Arthropoda Selecta 28(2): 277-290. full article (PDF). Reference page. .
• Omelko, M.M. & Omelko, N.V. 2019. Two New Species of the Gelechiid-Moth Genus Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae) from Laos. The perelegans and hibisci Species-Groups. Entomological Review 99(8): 1215-1221. (Russian Text © The Author(s), 2019, published in Zoologicheskii Zhurnal, 2019, 98(7): 785–791). DOI: 10.1134/S0013873819080165. . Reference page. 
• Omelko, M.M. & Omelko, N.V. 2019a. Two new species of Gelechiid moths genus Semophylax Meyrick, 1932 (Lepidoptera, Gelechiidae) found in Malaysia. Amurian zoological journal XI(1): 37-41. (in Russian). Reference page. 
• Omelko, M.M. & Omelko, N.V. 2019b. New genera and species of gelechiid moths from the subfamily Gelechiinae (Lepidoptera, Gelechiidae) from Laos and Malaysia. Amurian zoological journal XI(2): 131-140. (in Russian). Reference page. 

2020

• Fomichev, A.A. & Omelko, M.M. 2020. Three new species of the Acantholycosa-complex (Aranei: Lycosidae) from the south of the Russian Far East. Arthropoda Selecta 29(2): 262-272. DOI: 10.15298/arthsel.29.2.13. . Reference page. 
• Marusik, Y.M., Omelko, M.M. & Koponen, S. 2020. On two enigmatic spiders (Araneae: Cybaeidae & Phrurolithidae) from the Russian Far East. Zootaxa 4899(1): 247–258. DOI: 10.11646/zootaxa.4899.1.13. . Reference page. 
• Marusik, Y.M., Omelko, M.M., Simonov, P.S. & Mi, X.Q. 2020. A survey of Alenatea Song et Zhu, 1999 (Aranei: Araneidae). Arthropoda Selecta 29(4): 487–494. DOI: 10.15298/arthsel.29.4.12. .Full article PDF). Reference page. 
• Omelko, M.M. & Marusik, Yu.M. 2020. A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos. Raffles Bulletin of Zoology 68: 479-487. DOI: 10.26107/RBZ-2020-0066. . Reference page. 
• Omelko, M.M., Marusik, Yu.M. & Lyle, R. 2020. A survey of Diphya Nicolet, 1849 (Araneae: Tetragnathidae) from South Africa. Zootaxa 4899(1): 259–279. DOI: 10.11646/zootaxa.4899.1.14. . Reference page. 
• Omelko, M.M. & Omelko, N.V. 2020. Four New Species of Gelechiid Moths of the Genus Photodotis Meyrick, 1911 (Lepidoptera, Gelechiidae) from Borneo. Entomological Review 100(5): 706-713. (Original Russian Text published in Zoologicheskii Zhurnal, 2020, 99(4): 422–429). DOI: 10.1134/S0013873820050115. . Reference page. 
• Omelko, M.M. & Omelko, N.V. 2020a. A new genus of gelechiid moths of the tribe Litini (Lepidoptera, Gelechiidae: Gelechiinae) from Malaysia. Amurian Zoological Journal 12(3): 364–368. Reference page. 
• Omelko, M.M. & Omelko, N.V. 2020b. Two new species of Gelechiid moths (Lepidoptera, Gelechiidae) from the genera Deltophora Janse and Stegasta Meyrick from central Laos. Amurian zoological journal XII(1): 43-48. (in Russian). Reference page. 
• Omelko, M.M. & Omelko, N.V. 2020c. A new genus of Gelechiid moths of the tribe Litini (Lepidoptera, Gelechiidae: Gelechiinae) from Malaysia. Amurian zoological journal XII(3): 364–368. (in Russian). Reference page. 

2021

• Fomichev, A.A. & Omelko, M.M. 2021. A new species of the genus Bifurcia (Aranei: Linyphiidae) from primorsky krai, Russia. Far Eastern Entomologist 435: 20-24. DOI: 10.25221/fee.435.3. . Reference page. 
• Marusik, Yu.M. & Omelko, M.M. 2021. A new species and new records of Cybaeus L. Koch, 1868 (Aranei: Cybaeidae) in the Maritime Province of Russia. Arthropoda Selecta 30(2): 216–220. Reference page. 
• Omelko, M.M. & Marusik, Yu.M. 2021. New data on Linyphiidae spiders (Aranei) in the Maritime Province of Russia. Arthropoda Selecta 30(2): 245–256. Reference page. 
• Omelko, M.M. & Omelko, N.V. 2021. A New Genus and Five New Species of the Gelechiid-Moth Tribe Gelechiini (Lepidoptera: Gelechiidae, Gelechiinae) from Laos and Malaysia. Entomological Review 101(2): 247-255. (Original Russian Text published in Zoologicheskii Zhurnal, 2021, 100(3): 279-287). DOI: 10.1134/S0013873821020111. . Reference page. 
• Omelko, M.M. & Omelko, N.V. 2021a. New and interesting species of gelechiid moths of the genus Helcystogramma Zeller 1877 (Lepidoptera, Gelechiidae) from Laos and Malaysia Amurian zoological journal XIII(3): 418-422. (in Russian). Reference page. 
• Omelko, M.M. & Omelko, N.V. 2021b. New data on the fauna of gelechiid moths of the genus Photodotis Meyrick (Lepidoptera, Gelechiidae) from Borneo and the description of a new genus. Amurian zoological journal XIII(4): 460-466. (in Russian). Reference page. 
• Ponomarenko, M.G., Omelko, M.M. & Omelko, N.V. 2021. New genus of gelechiid moths (Lepidoptera: Gelechiidae) from Borneo, and its four new species. Zootaxa 5004(3): 465–480. DOI: 10.11646/zootaxa.5004.3.4. . Reference page. 
• Ponomarenko, M.G., Omelko, M.M. & Omelko, N.V. 2021a. Neolitinia sulphurata gen. n., sp. n. (Lepidoptera: Gelechiidae) from Borneo Island. Far Eastern Entomologist 438: 6-12. DOI: 10.25221/fee.438.2. . Reference page. 

2022

• Omelko, M.M. & Marusik, Yu.M. 2022. A new species of the genus Borboropactus Simon, 1884 (Aranei: Thomisidae) from Borneo Island, Malaysia. Far Eastern Entomologist 459: 27-32. DOI: 10.25221/fee.459.2 . Full article (PDF). Reference page. 
• Ponomarenko, M.G., Omelko, M.M. & Omelko, N.V. 2022. Parateleiopsis feregrisea (Lepidoptera : Gelechiidae), a new genus and a new species from Borneo. Annales Zoologici Fennici 59(1): 47-52. DOI: 10.5735/086.059.0105. . Reference page. 

2023

• Fomichev, A.A. & Omelko, M.M. & Marusik, Yu.M. 2023. A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353(2): 117–130. DOI: 10.11646/zootaxa.5353.2.2. Reference page. 
• Omelko, M.M. 2023. Two new species of Kiekie Polotow & Brescovit, 2018 (Araneae: Ctenidae) from the highlands of Panama. Zootaxa 5323(2): 275–284. DOI: 10.11646/zootaxa.5323.2.7. Reference page. 
• Omelko, M.M. & Fomichev, A.A. 2023. A survey of Sumatran Ctenidae (Araneae). 2. Three new species of Bowie Jäger, 2022. Zootaxa 5353(2): 101–116. DOI: 10.11646/zootaxa.5353.2.1. Reference page. 
• Omelko, M.M. & Fomichev, A.A. 2023. Two species of the genus Psechrus Thorell, 1878 (Araneae: Psechridae) from North Sumatra, Indonesia. Far Eastern Entomologist 482: 1–9. DOI: 10.25221/fee.482.1 Reference page. 
• Omelko, M.M. & Omelko, N.V. 2023. Two New Species of the Gelechiid-Moth Genus Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae) from Panama Entomological Review 103(2): 241–246. (Russian Text © The Author(s), 2023, published in Zoologicheskii Zhurnal, 102(3): 304–309). DOI: 10.1134/S0013873823020112. . Reference page. 
• Ponomarenko, M.G., Omelko, M.M. & Omelko, N.V. 2023. New species of gelechiid moths of the genus Sabaha M. Omelko et N. Omelko, 2019 (Lepidoptera: Gelechiidae) from Borneo Island. Far Eastern Entomologist 486: 6-24. DOI: 10.25221/fee.486.2. . Reference page. 

2024

• Omelko, M.M. & Fomichev, A.A. 2024. A survey of the Sumatran Ctenidae (Araneae). 3. Amauropelma mariae sp. n., one of the tiniest species of the genus. Zootaxa 5474(2): 173–180. DOI: 10.11646/zootaxa.5474.2.5. . Reference page. 
• Ponomarenko, M.G. & Omelko, M.M. 2024. A new genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae, Gelechiinae) from the East Malaysia with unusual male secondary characters. Zootaxa 5447(4): 573–582. DOI: 10.11646/zootaxa.5447.4.8. . Reference page.