M. Alex Smith

M. Alex Smith, Canadian entomologist

• Department of Integrative Biology, University of Guelph, Guelph, ON, Canada.

Taxon names authored

(List may be incomplete)

• 14 taxon names authored by M. Alex Smith

Eponyms

(List may be incomplete)

• 2 eponyms of M. Alex Smith

Publications

(List may be incomplete)

2011

• Sharkey, M.J., Clutts, S., Tucker, E.M., Janzen, D., Hallwachs, W., Dapkey, T. & Smith, M.A. 2011. Lytopylus Förster (Hymenoptera, Braconidae, Agathidinae) species from Costa Rica, with an emphasis on specimens reared from caterpillars in Area de Conservación Guanacaste. ZooKeys 130: 379–419. DOI: 10.3897/zookeys.130.1569 . Reference page. 

2012

• Butcher, B.A., Smith, M.A., Sharkey, M.J. & Quicke, D.L.J. 2012. A turbo-taxonomic study of Thai Aleiodes (Aleiodes) and Aleiodes (Arcaleiodes) (Hymenoptera: Braconidae: Rogadinae) based largely on COI barcoded specimens, with rapid descriptions of 179 new species. Zootaxa 3457: 1–232. Preview ISBN 978-1-77557-000-4 (paperback) ISBN 978-1-77557-001-1 (Online edition) Reference page. 
• Whitfield, J.B., Fernández-Triana, J.L., Janzen, D.H., Hallwachs, W, Smith, M.A. & Cardinal, S. 2012. Mariapanteles (Hymenoptera, Braconidae), a new genus of Neotropical microgastrine parasitoid wasp discovered through biodiversity inventory. ZooKeys 208: 61-80. DOI: 10.3897/zookeys.208.3326 . Reference page. 
• López-Estrada, E.K., Briceño G., R.A., Smith, M.A., Nunes, J., Penteado-Dias, A.M., Ceccarelli, F.S., Clebsch, H. & Zaldivar-Riverón, A. 2012. Seven new species of Notiospathius (Hymenoptera, Braconidae, Doryctinae) from Northwest Venezuela. Journal of Hymenoptera Research. 29:37-61. DOI: 10.3897/jhr.29.3555. Reference page. 
• Quicke, D.L.J., Smith, M.A., Janzen, D.H., Hallwachs, W., Fernandez-Triana, J., Laurenne, N.M., Zaldívar-Riverón, A., Shaw, M.R., Broad, G.R., Klopfstein, S., Shaw, S.R., Hrcek, J., Hebert, P.D.N., Miller, S.E., Rodriguez, J.J., Whitfield, J.B., Sharkey, M.J., Sharanowski, B.J., Jussila, R., Gauld, I.D., Chesters, D. & Vogler, A.P. 2012. Utility of the DNA barcoding gene fragment for parasitic wasp phylogeny (Hymenoptera: Ichneumonoidea): data release and new measure of taxonomic congruence. Molecular Ecology Resources 12: 676–685. DOI: 10.1111/j.1755-0998.2012.03143.x Reference page. 

2014

• Fernández-Triana, J., Janzen, D., Hallwachs, W., Whitfield, J.B., Smith, M.A. & Kula, R. 2014. Revision of the genus Pseudapanteles (Hymenoptera, Braconidae, Microgastrinae), with emphasis on the species in Area de Conservación Guanacaste, northwestern Costa Rica. ZooKeys, 446: 1–82. DOI: 10.3897/zookeys.446.8195 . Reference page. 
• Fernandez-triana, J.L., Whitfield, J.B., Smith, M.A., Hallwachs, W. & Janzen, D.H. 2014. First record of the genus Venanus (Hymenoptera: Braconidae: Microgastrinae) in Mesoamerica, with the description of two new species from Costa Rica. Biodiversity Data Journal 2: e4167. DOI: 10.3897/BDJ.2.e4167 . Reference page. 

2015

• Fernandez-Triana, J.L., Whitfield, J.B., Smith, M.A., Kula, R.R., Hallwachs, W. & Janzen, D.H. 2015. Revision of the genera Microplitis and Snellenius (Hymenoptera, Braconidae, Microgastrinae) from Area de Conservacion Guanacaste, Costa Rica, with a key to all species previously described from Mesoamerica. Deutsche Entomologische Zeitschrift 62(2): 137–201. DOI: 10.3897/dez.62.5276. Reference page. 
• Fleming, A.J., Wood, D.M., Janzen, D.H., Hallwachs, W. & Smith, M.A. 2015. Seven new species of Spathidexia Townsend (Diptera: Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, Costa Rica. Biodiversity Data Journal 3: e4597. DOI: 10.3897/BDJ.3.e4597. Reference page. 
• Fleming, A.J., Wood, D.M., Smith, M.A., Hallwachs, W. & Janzen, D. 2015. Three new species of Ametadoria Townsend (Diptera: Tachinidae) from Area de Conservación Guanacaste, Costa Rica. Biodiversity Data Journal 3: e5039. DOI: 10.3897/BDJ.3.e5039 . Reference page. 
• Fleming, A.J., Wood, D.M., Janzen, D.H., Hallwachs, W. & Smith, M.A. 2015. Three new species of Trigonospila Pokorny (Diptera: Tachinidae), from Area de Conservación Guanacaste, northwestern Costa Rica, with a key for their identification. Biodiversity Data Journal 3: e4595. DOI: 10.3897/BDJ.3.e4595. Reference page. 
• Fleming, A.J., Wood, D.M., Janzen, D.H., Hallwachs, W. & Smith, M.A. 2015. Corrigendum: Three new species of Trigonospila Pokorny (Diptera: Tachinidae), from Area de Conservación Guanacaste, northwestern Costa Rica, with a key for their identification. Biodiversity Data Journal 3: e6459. DOI: 10.3897/BDJ.3.e6459. Reference page. 
• Fleming, A.J., Wood, D.M., Smith, M.A., Janzen, D.H. & Hallwachs, W. 2015. Nine new species of Itaplectops (Diptera: Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, northwestern Costa Rica, with a key to Itaplectops species. Biodiversity Data Journal 3: e4596. DOI: 10.3897/BDJ.3.e4596. Reference page. 
• Hansson, C., Smith, M.A., Janzen, D.H. & Hallwachs, W. 2015. Integrative taxonomy of New World Euplectrus Westwood (Hymenoptera, Eulophidae), with focus on 55 new species from Area de Conservación Guanacaste, northwestern Costa Rica. ZooKeys, 485: 1–236. DOI: 10.3897/zookeys.458.9124Reference page. 
• Sharkey, M.J., Chapman, E.G., Janzen, D.H., Hallwachs, W. & Smith, M.A. 2015. Revision of Aphelagathis (Hymenoptera, Braconidae, Agathidinae, Agathidini). Zootaxa 4000(1): 73–89. DOI: 10.11646/zootaxa.4000.1.3. Preview (PDF) Reference page. 

2016

• Fernandez-Triana, J.L., Whitfield, J.B., Smith, M.A., Dapkey, T., Hallwachs, W. & Janzen, D.M. 2016. Review of the world species of Exoryza (Hymenoptera, Braconidae, Microgastrinae), with description of five new species. Deutsche Entomologische Zeitschrift 63(2): 195–210. DOI: 10.3897/dez.63.8977. Reference page. 
• Fernandez-Triana, J.L., Boudreault, C., Dapkey, T., Smith, M.A., Rodriguez, J., Hallwachs, W. & Janzen, D.H. 2016. Revision of the genus Promicrogaster (Hymenoptera, Braconidae, Microgastrinae) from Area de Conservación Guanacaste, Costa Rica, with a key to all species previously described from Mesoamerica. Journal of Hymenoptera Research 50: 25–79. DOI: 10.3897/JHR.50.8220. Reference page. 
• Fleming, A.J., Wood, D.M., Smith, M.A., Janzen, D.H., Hallwachs, W. & Dapkey, T. 2016. A new species of Phosocephala Townsend, 1908 (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica. Biodiversity data journal 4: e7863. DOI: 10.3897/BDJ.4.e7863 Reference page. 
• Fleming, A.J., Wood, D.M., Smith, M.A., Hallwachs, W., Janzen, D. & Dapkey, T. 2016. Two new species of Erythromelana Townsend, 1919 (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica. Biodiversity Data Journal 4: e7386. DOI: 10.3897/BDJ.4.e7386. Reference page. 
• Quicke, D.L.J., Belokobylskij, S.A., Smith, M.A., Rota, J., Hrcek, J. & Butcher, B.A. 2016. A new genus of rhysipoline wasp (Hymemoptera: Braconidae) with modified wing venation from Africa and Papua New Guinea, parasitoid on Choreutidae (Lepidoptera). Annales Zoologici 66: 173–192. DOI: 10.3161/00034541ANZ2016.66.2.003 Reference page. 

2017

• Fleming, A.J., Wood, D.M., Smith, M.A., Dapkey, T., Hallwachs, W. & Janzen, D.H. 2017. Five new species of Vibrissina Rondani (Diptera: Tachinidae) from Area de Conservación Guanacaste in Northwestern Costa Rica. Biodiversity Data Journal 5: e10967. DOI: 10.3897/BDJ.5.e10967. Reference page. 
• Fleming, A.J., Wood, D.M., Smith, M.A., Hallwachs, W., Janzen, D. & Dapkey, T. 2017. Nine new species of Uramya Robineau-Desvoidy (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica, with a key to their identification. Biodiversity Data Journal 5: e9649. DOI: 10.3897/BDJ.5.e9649. Reference page. 
• Fleming, A.J., Wood, D.M., Smith, M.A., Dapkey, T., Hallwachs, W. & Janzen, D. 2017. A new species of Voria Robineau-Desvoidy (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica. Biodiversity Data Journal 5: e20123. DOI: 10.3897/BDJ.5.e20123. Reference page. 
• Kang, I., Chapman, E.G., Janzen, D.H., Hallwachs, W., Dapkey, T., Smith, M.A. & Sharkey, M.J. 2017. Revision of the species of Lytopylus from Area de Conservación Guanacaste, northwestern Costa Rica (Hymenoptera, Braconidae, Agathidinae). ZooKeys 721: 93–158. DOI: 10.3897/zookeys.721.20287. Reference page. 

2018

• Fleming, A.J., Wood, D.M., Smith, M.A., Hallwachs, W. & Janzen, D.H. 2018. Revision of the Mesoamerican species of Calolydella Townsend (Diptera: Tachinidae) and description of twenty-three new species reared from caterpillars in Area de Conservación Guanacaste, northwestern Costa Rica. Biodiversity Data Journal 6: e11223. DOI: 10.3897/BDJ.6.e11223. Reference page. 

2019

• Arias-Penna, D., Whitfield, J.B., Janzen, D.H., Hallwachs, W., Dyer, L.A., Smith, M.A., Hebert, P.D.N. & Fernández-Triana, J.L. 2019. A species-level taxonomic review and host associations of Glyptapanteles (Hymenoptera, Braconidae, Microgastrinae) with an emphasis on 136 new reared species from Costa Rica and Ecuador. ZooKeys, 890: 1–685. DOI: 10.3897/zookeys.890.35786 Reference page. 
• Fernandez-Triana, J.L., Boudreault, C., Dapkey, T., Smith, M.A., Hallwachs, W. & Janzen, D.H. 2019. A revision of Dolichogenidea (Hymenoptera, Braconidae, Microgastrinae) with the second mediotergite broadly rectangular from Area de Conservación Guanacaste, Costa Rica. Zookeys, 835: 87–123. DOI: 10.3897/zookeys.835.33440 Reference page. 
• Fleming, A.J., Wood, D.M., Smith, M.A., Dapkey, T., Hallwachs, W. & Janzen, D. 2019. A new species of Trismegistomya Reinhard (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica. Biodiversity Data Journal, 7: e29130. DOI: 10.3897/BDJ.7.e29130 Reference page. 
• Fleming, A.J., Wood, D.M., Smith, M.A., Dapkey, T., Hallwachs, W. & Janzen, D.H. 2019. Twenty-two new species in the genus Hyphantrophaga Townsend (Diptera: Tachinidae) from Area de Conservación Guanacaste, with a key to the species of Mesoamerica. Biodiversity Data Journal, 7: e29553. DOI: 10.3897/BDJ.7.e29553 Reference page. 
• Meierotto, S., Sharkey, M.J., Janzen, D.H., Hallwachs, W., Hebert, P.D.N., Chapman, E.G. & Smith, M.A. 2019. A revolutionary protocol to describe understudied hyperdiverse taxa and overcome the taxonomic impediment. Deutsche Entomologische Zeitschrift 66: 119–145. DOI: 10.3897/dez.66.34683 . Reference page. 
• Sharkey, M.J., Penteado-Dias, A.M., Smith, M.A., Hallwachs, W. & Janzen, D. 2019. Synopsis of New World Sigalphinae (Hymenoptera, Braconidae) with the description of two new species and a key to genera. Journal of Hymenoptera Research, 68: 1–11. DOI: 10.3897/jhr.68.30131 Reference page. 

2021

• Quicke, D.L.J., Fagan-Jeffries, E.P., Jasso-Martínez, J.M., Zaldívar-Riverón, A., Shaw, M.R. & Janzen, D.H., Hallwachs, W., Smith, M.A., Hebert, P.D.N., Hrcek, J., Miller, S.E., Sharkey, M.J., Shaw, S.R. & Butcher, B.A. 2021. A molecular phylogeny of the parasitoid wasp subfamily Rogadinae (Ichneumonoidea: Braconidae) with descriptions of three new genera. Systematic Entomology 46: 1019–1044. DOI: 10.1111/syen.12507 Reference page. 
• Sharkey, M.J., Baker, A., McCluskey, K., Smith, A., Naik, S., Ratnasingham, S., Manjunath, R., Perez, K., Sones, J., D'Souza, M., St. Jacques, B., Hebert, P., Hallwachs, W. & Janzen, D. 2021. Addendum to a minimalist revision of Costa Rican Braconidae: 28 new species and 23 host records. ZooKeys 1075ː 77–136. DOI: 10.3897/zookeys.1075.72197 Reference page. 
• Sharkey, M.J., Janzen, D.H., Hallwachs, W., Chapman, E.G., Smith, M.A., Dapkey, T., Brown, A., Ratnasingham, S., Naik, S., Manjunath, R., Perez, K., Miltom, M., Hebert, P., Shaw, S.R., Kittel, R.N., Solis, M.A., Metz, M.A., Goldstein, P.Z., Brown, J.W., Quicke, D.L.J., van Achterberg, C., Brown, B.V. & Burns, J.M. 2021. Minimalist revision and description of 403 new species in 11 subfamilies of Costa Rican braconid parasitoid wasps, including host records for 219 species. ZooKeys 1013: 1–665. DOI: 10.3897/zookeys.1013.55600. . Reference page. 

2023

• Fernández-Triana, J.L., Shimbori, E.M., Whitfield, J.B., Penteado-Dias, A.M., Shaw, S.R., Boudreault, C., Sones, J., Perez, K., Brown, A., Manjunath, R., Burns, J.M., Hebert, P.D.N., Smith, M.A., Hallwachs, W. & Janzen, D.H. 2023. A revision of the parasitoid wasp genus Alphomelon Mason with the description of 30 new species (Hymenoptera, Braconidae). Zookeys 1175ː 5–162. DOI: 10.3897/zookeys.1175.105068 Reference page. 

2025

• Fernandez-Triana, J.L., Boudreault, C., Whitfield, J.B., Höcherl, A., Smith, M.A., Hallwachs, W. & Janzen, D.H. 2025. A revision of the parasitoid wasp genus Dolichogenidea Viereck (Hymenoptera, Braconidae) in the Neotropical region, with the description of 102 new species. ZooKeys 1237: 1–250. DOI: 10.3897/zookeys.1237.141007 Reference page.