User:Circeus/article sandbox

[this preprint was originally written on the author's Wikispecies' userspace]

In integrating data from the recently published catalogue of Thysanopteran genera (Mound & Hastenpflug-Vesmanis, 2021, which I will refer to as the Catalogue throughout for simplicity) in the Internet Register of Marine and Nonmarine Genera (IRMNG, Rees & al., 2020), a number of issue of a nomenclatural or bibliographic nature were noted and compiled. Following private communications with the primary author of that catalogue, the following addenda and corrigenda were prepared.

Workflow and nature of errors revealedEdit

The first step consists of verifying the name's existence in IRMNG through basic, then fuzzy search. The latter serves to detect potential misspellings or variants, such as those generated by ligatures (i.e. æ and œ). A significant number of these misspellings were documented by or errors introduced in the original Nomenclator Zoologicus, a few additional ones (e.g. "Australaealles Rheinheimer, 1993") were further generated in the transition to its online database form (Remsen et al., 2016). The former detects potential homonyms or duplicate records. In groups such as Coleoptera many names are listed under conflicting authorships because of diverging presentation or opinion in imported databases. The duplicate entries si generated must be cleaned up and properly synonymized. This was generally not the case within Thysanoptera, although a few misspelling were noticed.

The second key step is entering the reference in IRMNG for use as Original Description source within the Aphia software (Vandepitte et al., 2015). This step often reveals a second set of errors. These include references that are incorrectly cross-referenced, ambiguous because they are under-differentiated, or missing entirely. A more subtle type of error is the case where a name is cited or spelled differently as a name author and as an article author. This problem is probably most commonly encountered with authors bearing compound Spanish or Portuguese family names (Kury et al., 2020:7-12).

The next step consist in a thorough online search to check for online copies of the original published material. Such material may be found on Google Books (in full or snippet form) or journal websites as well as a variety of institutional or independent repositories, including websites such as the Internet Archive, ResearchGate, the Biodiversity Heritage Library and Zobodat. While this third step may be considered a mere adjunct to the previous step, it is where a different set of issues often come to the fore: incorrect or misleading bibliographical details. Such error include the common issue of bibliographical vs. printing date, but also page, issue or volume numbers as well as malformed or incomplete serial titles. The omission of series information required to correctly locate a reference is a distressingly common across taxonomic literature.

At the fourth step, original material will be double-checked if available. This is primarily a verification that the name is indeed made available there, but also ensures no other Code-related issues are present. Such issues include replaced names not otherwise noted and incorrect spellings both original or subsequent. It also constitutes a last line of defense against two references being confused with each other. In entering a previous catalogue (Francke, 2019), an error where two pinpoint references differed only by the issue number (with year, author and page number all otherwise identical and legitimate) would not have been detected but for this step.

Finally, once all catalogued names in given taxonomic group has been fully handled (either an order or a family), any name not yet treated is examined. If possible, it is checked against the literature in order to verify its taxonomic and nomenclatural status. This final step is where leftover misspellings or variants are correctly synonymized, leaving only the more obscure overlooked names. Such leftover name are then handled as much as can be from literature available online.

Because IRMNG data is imported in other databases such as GBIF, it is crucial to prevent erroneous data from entering it and then spreading to the point it becomes embedded as valid data despite being nonexistent in the literature. This workflow is implemented with that goal in mind, but it is also uniquely suited to revealing erroneous information that may appear in a catalogue being entered.

An obvious shortcoming of this process is that it is, by the very nature of IRMNG, primarily concentrated on genus-group names. Generally, no effort was made to verify that the bibliographical or nomenclatural accuracy of other data (e.g. that related to type species and synonymization sources). Such issues are noted only occasionally as they were noticed while in the process of verifying other details. An exception was made for newly added entries, for which every attempt was made to provide as complete information as was given for existing entries.

Although cognizant in the workings of the International Code of Zoological Nomenclature, the author is not himself a taxonomist and lacks access to robust interlibrary loan systems to consult more obscure literature. This means that in several cases, the current status of names documented in IRMNG but not in the Catalogue could not be reliably assessed.


Dating and formatting of referencesEdit

The publication dates found on the work were generally assumed to be accurate. Where explanation was felt pertinent, it was included in the relevant section. Where that date is different from the formal year as given—such as for yearly-dated serials—,the correct date is given in parentheses, and the date which would be otherwise cited is given in brackets at the end of the citation.

New references were generally inserted so as to avoid disrupting the references present whenever possible rather than in any attempt at keeping them chronologically ordered within each year. Existing references years only had differentiating letters added where new references were added or existing references lacked such needed differentiation. Otherwise newly added reference where placed sequentially after the existing one. References required by new material added here are noted directly alongside the relevant material. If a reference is merely missing, it is listed directly in the "Missing references" section. No references are listed as being deleted.

To simplify cross-referencing, references also found in Mound & Hatenpflug (2021) are cited with the same year disambiguation as in that paper regardless of the need for disambiguation here. The only exceptions are years emended here for the purpose of newly needed disambiguation.

Issue numbers, in so far as they could be determined, are included for all references added to the original article. Issue numbers are only noted for reference emendation if that part of the reference is already being corrected or where issues were found to be separately paginated.

Ordering and formatting of entriesEdit

While addenda and corrigenda of this nature (e.g. Bouchard & Bousquet, 2020; Lyal & Alonso-Zarazaga, 2006; Alonso-Zarazaga et al., 2016) are traditionally structured in a strict sequential format, it was felt somewhat more convenient to structure this contribution in a thematic fashion by grouping corrections according to their nature. Page number were nonetheless included for each individual emendation, except those concerning added or emended references.

Entries have been ordered in roughly decreasing order of their taxonomic and nomenclatural significance. Starting with names omitted entirely and dates to be corrected, then authorships to be emended. Following this are other nonbibliographic emendations such as adjustments to reflect technical nomenclatural terms or mentions of earlier homonyms. Finally, three sections correct erroneous cross-references, add missing references, and emend erroneous bibliographical data in the reference section. Within each section, entries are presented sequentially as they appear in the paper.

References whose emendation affect a name's authorship or dating are emended or replaced alongside the entry for the name itself. Those changes are not listed in the sectoins of the article that deal solely with references.

Entries for synonyms are treated separately from their valid name, except where the addition of a new subentry requires changing the form of the main entry itself. Notably, the Catalogue is not consistent in its treatment of replaced homonyms. Some of these names are listed separately, some are combined into the entry for the replacement name and other are missing entirely. The first of these three options is here applied consistently across the board.


Names of indeterminate statusEdit

The following names are documented in IRMNG, but lacked mentions in subsequent literature. Their status and typification are unknown.

Achaetothrips Shumsher, 1946: 157,161. Junior homonym of Achaetothrips Karny, 1908.

Allucita Hill, 1775: :21

This extremely obscure name and the two species placed under it are definitely nomina oblita, but they must be synonymized before reversal of precedence under art. 23.9 can apply.

Glabrothrips Velikan, 1986: 720.

Macroplatothrips and Rostrothrips Duan in Wu & Pan (2001: 176)

  • Wu, H. & Pan, C. (2001) Insects of Tianmushan National Nature Reserve. Science Press, Beijing. 764 pp. ISBN 978-7-03-009871-9.

It was not possible to determine with any likelihood which of Ereikithrips or Ereikethrips was used by Knechtel (1960).

Missing entries and nomenclatural issuesEdit

Page 6: Three names must be added to the synonymy of Heterothrips Hood. The synonymy author for Protemnothrips is unknown but must postdate Jacot-Guillamod (1970).

Phyllothrips Buffa, 1908c. Type species Phyllothrips pilosus Buffa, 1908c, by monotypy. Synonymy by Bagnall (1910d).
Phyllothripsiolus Strand, 1912. Replacement name for Phyllothrips Buffa, 1908c not Phyllothrips Hood, 1908b.
Heterothrips (Protemnothrips) Hood, 1937a: 258. Type species: Heterothrips miconiae, by monotypy. Synonymy by ???.

Page 7 (1): Liassothrips is a replacement name, the entry should be emended as follow:

Liassothrips Priesner, 1949b [1 species]. Replacement name for Mesothrips Martynov, 1927, not Mesothrips Zimmermann, 1900.
Mesothrips Martynov, 1927. Type species Mesothrips crassipes Martynov, 1927, by monotypy.
  • Martynov, A.V. (1927). [Jurassic fossil insects from Turkestan]. Bulletin of the Academy of Sciences USSR 7: 757-768. [in Russian]

Page 7 (2): Under Archankothrips, add the following overlooked synonym. No source could be found originating this synonymy, but it predates Ulitzka (2017).

Promelanthrips Priesner, 1929a: 112. Type species Promelanthrips spiniger Priesner, 1929, by monotypy. Synonymy by ???.

Page 12: The following available emendation should be added as a synonym of Aptinothrips:

Aptenothrips Agassiz, 1846. Unjustified emendation of Aptinothrips Haliday, 1836.
  • Agassiz, L. (1846). Nomenclatoris zoologici. Index universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum, nec non variis adnotationibus et emendationibus. Jent et Gassmann, Soloduri. viii + 393 pp.

Page 16: The situation surrounding the correct type of Frankliniella is complex and merits discussion. When coining the name, Karny (1910) explicitly did so as a new replacement name for Physapus Amyot & Serville, 1843. Referring to it as "Serville's name", he states: "I have thus used Serville's name in this sense in order not to have to introduce a new name into the literature."[1] This represents a problem because article 67.8 states that a replacement name must by definition have the type of the name it replaces i.e.Physapus Amyot & Serville, 1843.[2] This makes the designation by Hood (1914b) invalid under the code.

Instead we must look for the valid type of Physapus Amyot & Serville. That name is omitted from the catalogue, but as far as could be determined, it was typified by Karny (1907:45) with "Physapus ater" Degeer, 1773. DeGeer (1773) does not actually use binomials for his thrips, or even the name Physapus, conceding to Linnaeus on the use of Thrips. For an explanation of how to interpret DeGeer's names, see Welter-Schultes & Wieland (2012). Karny does, however, synonymize this name with Thrips vulgatissimus Haliday, 1836 which we must take as being designated as type. This is clearly a problem as T. vulgatissimus is a species in good standing of Thrips Linnaeus, 1758. Its acceptance as type would be incredibly disruptive, causing Frankliniella to fall in synonymy with Thrips and the obscure Exophthalmothrips Moulton, 1933a to become the valid generic name for over a fifth of all species currently accepted in Thysanopera. Only a request for conservation would prevent this infelicitous result were it not for another important factor.

In both publications, Karny (1907, 1910) explicitly mentions that he is following Uzel's concepts. As noted by Hood (1914b), Uzel's concept of T. vulgatissimus actually corresponds to Thrips intonsa Trybom, 1895 (cf. Uzel, 1895:455-456). This situation is here resolved fully by formally acknowledging this misidentification and designating here (under Article 70.3.2 of the Code) the type species of Physapus Amyot & Serville, 1843:643 as Thrips intonsa Trybom, 1895, misidentified as Thrips vulgatissimus Haliday, 1836 in the subsequent designation by Karny (1907).

Following the above discussion, the entry for Frankliniella must be emended as follow:

Frankliniella Karny, 1910 [236 species]. Replacement name for Physapus Amyot & Serville, 1843 not Physapus Leach, 1815.
Physapus Amyot & Serville, 1843. Type species Thrips intonsa Trybom, 1895, misidentified as Thrips vulgatissimus Haliday, 1836 in the subsequent designation by Karny (1907).

Page 17: The name Ctenothripiella has been more or less unviersally associated with Priesner (1927). It is not clear whether Priesner's name is actually available under Art. 11.6.1. of the code. This is however irrelevant as the name was in fact published the year before by Moulton as a subgenus of Thrips. Furthermore, although the name is spelled by modern authors as Ctenothripiella, Priesner and his contemporary consistently used the spelling Ctenothripella,[3] with a single i. The entry must be emended as follow:

Thrips (Ctenothripella) Moulton, 1926b. Type species Thrips gillettei Moulton, 1926, by monotypy. Synonymy by Priesner (1927).

Page 22: Under Thrips, add the following synonym:

Physopus Rafinesque, 1815. Unnecessary replacement name for Thrips Linnaeus, 1758. Synonymy by Gentile & Bailey (1968).
  • Gentile, A.G. & Bailey, S.F. (1968). A revision of the genus Thrips Linnaeus in the New World, with a catalogue of world species (Thysanoptera: Thripidae). University of California publications in entomology, 51: 1-95.
  • Rafinesque, C.S. (1815). Analyse de la nature, ou tableau de l'universe et des corps organisés. Palermo: Jean Barravecchia. 224 pp.

Page 23: The correct date and place of publication of Euthrips is Targioni-Tozzettti (1881). Targioni-Tozzetti (1887) is the place of publication of Thrips (Euthrips) consociatus and is at best a later usage of the name. Although Karny [1910] took the name as available from that publication, this version of Euthrips would be a junior synonym of Mycterothrips. There is no typification as Hood (1914b) explained the name to be an unnecessary replacement. The entry should be amended as follow:

Euthrips Targioni-Tozzettti, 1881. Unnecessary replacement name for Thrips Linnaeus, 1758. Synonymy by Hood (1914b).
  • Targioni-Tozzetti, A. (1881). Relazione intorno ai lavori della R. Stazione di Entomologia Agraria di Firenze per gli anni 1877–78. Annali di Agricoltura. 34: 134-161.

Page 30: Because "Ahamothrips" is not an available name, it cannot by definition be valid and it should probably not be listed in bold as though it were. The following emendation is suggested:

"Ahamothrips" Kumar, Tyagi & Bhatti, 2007 [1 species]. Unavailable name: no type specimen was designated for the type species "Ahamothrips maxima".

Page 32: A similar emendation is recommended for "Chalepothrips":

"Chalepothrips" Priesner, 1968 [1 species]. Unavailable name: the purported type species "Chalepothrips ardisiae" was not described or made available.

Page 34: Under Gigantothrips, add the following overlooked synonym:

Cercothrips Hood, 1919a:73. Type species Acanthinothrips nigrodentatus Karny by original designation. Synonymy by Mound (1968a).

Page 35: Under Glyptothrips, add the synonym:

Pactothrips Hood, 1912d. Unavailable name: incorrect original spelling for Glyptothrips.

Page 37-38: Some clarification is needed to the entries for Karnyothrips and the various homonyms in its synonymy. Most noticeable is a missing entry for Karnyia Watson, 1922. The entries for Karnyothrips, Karynia and Watsoniella should be replaced by the following:

Karnyothrips Watson, 1923 [49 species]. Replacement name for Karnyia Watson, 1922 not Karnyia Shelford [Blattodea].
Karnyia Watson, 1922. Type species Karnyia weigeli Watson, 1922 (=Anthothrips flavipes Jones, 1912), by monotypy.
Karynia Watson, 1922. Incorrect original spelling for Karnyia Watson.
Dolichothrips Watson, 1920. Type species Cephalothrips elongatus Watson, 1919 (=Karnyothrips melaleucus Bagnall), by monotypy. Synonymy by Stannard (1957).
Watsoniella Karny, 1923. Replacement name for Dolichothrips Watson, 1920 not Dolichothrips Karny, 1912. Junior homonym of Watsoniella Berg, 1898 [Lepidoptera, Hesperidae].

Page 38: Insert the following entry after Kellyia:

Kilkuyrukus Özdikmen, 2008 [1 species]. Replacement name for Schlechtendalia Bagnall, 1929b not Schlechtendalia Lichtenstein, 1883 [Hemiptera, Aphididae].
Schlechtendalia Bagnall, 1929b. Type species Schlechtendalia longituba Bagnall, 1929b, by monotypy.
  • Özdikmen, H. (2008). A new name, Kilkuyrukus for the preoccupied thrips genus Schlechtendalia Bagnall, 1929 (Thysanoptera). Munis Entomology & Zoology. 3(2):767-768.

Page 41 (1): Under the code, the names Phaenothrips Priesner, 1968 and Phenothrips Ananthakrishnan, 1968 are not homonyms. The priority of the former over the replacement name Phenicothrips must be restored:

Phaenothrips Priesner, 1968a [8 species]. Type species Gynaikothrips daetymon Karny, 1923, by original designation.
Phenicothrips Bhatti, 1995c. Unnecessary replacement name for Phaenothrips Priesner [Phaenothrips Priesner and Phenothrips Ananthakrishnan, 1968 are not homonyms under Art. 56.1. Art. 58.1 does not apply to genus-group names.]

Page 41 (2): there is no evidence that Burmeister (1838) specifically intended to change the spelling of Haliday's Phlaeothrips nor that Amyot & Serville are taking his spelling over Haliday's. In those circumstances (ICZN Article 33.5), the name must be considered an incorrect spelling:

Phloeothrips Burmeister, 1838. Incorrect subsequent spelling of Phlaeothrips Haliday, 1836.

Page 41 (3): As noted earlier under Frankliniella, Euphlaeothrips Morison, being a replacement name for a misinterpretation, has no type designation of its own:

Euphlaeothrips Morison, 1949. Unnecessary replacement name for Phlaeothrips Haliday [as "Phloeothrips Amyot & Serville"].

Page 45: Under Trybomia, add the following overlooked synonym. The attribution for synonymy is taken from Retana-Salazar (2015) as Moulton's publication could not be seen.

Ommatothrips Hood, 1915c. Type species Ommatothrips gossypii Hood, 1915c , by monotypy. Synonymy by Moulton (1933).

Authorships or dates to be correctedEdit

Page 5: The date for Aeolothrips (Gevarothrips) Lacasa Plasencia, 1983 must be correct to 1984. Volume 59 of Eos correspond to publication year 1983, but is internally dated to March 1984. Furthermore, the corresponding reference was given under the author name "Plasencia" instead of "Lacasa Plasencia". It should emended as follow:

  • Lacasa Plasencia, A. (1984). Nuevo Tisanoptero de la Península Ibérica, Aeolothrips bournieri sp.n. (Aeolothripidae). Eos. 59(1–4): 67–75 [1983].

Page 6: The correct place of publication for Palaeothrips is not Scudder (1875b), but rather (as cited in that work) Scudder (1868:221). The full reference is:

Page 13: To correctly match its reference, the authorship of Brooksithrips should be corrected from Retana & Mound to Retana-Salazar & Mound.

Page 14: The date for Cricothrips should be corrected from 1913 to 1912c, and the reference "Trybom (1913)" emended accordingly in the references. At the bottom of page 52 of Trybom (1912c) is a printing date ("tryckt den") of November 6, 1912. Although the 1913 date is also given in Nomenclator Zoologicus, there does not seem to be any clear reason for disbelieving the 1912 date.

Page 15: The name Eremiothrips is consistently referred to as published on page 44 of Priesner (1950a). However, that article only runs to page 37. Page 44 falls within Priesner (1950c). Nomenclator Zoologicus gives the pages number as the more likely 28. Page 44 could be referring to "Priesner (1949b)" of the Catalogue, but this only further muddies the situation. Because the relevant volumes of the Bulletin de la Société Royale entomologique d'Égypte are not available online, it was not possible to confirm the exact place and year of publication of this genus between Priesner (1949b, 1950a, 1950c).

Page 17: The name Parahysopus was published twice by Girault in 1927. Once in Girault (1927), and once in an issue (issue four of volume 14) of Insecutor Inscitiae Menstruus with a pritned date of April 26, 1927. It was not possible to determine which of these was issued earliest.

  • Girault, A.A. (1927) Notes on Australian Thysanoptera. Insecutor Inscitiae Menstruus 14(4): 188-190

Page 19 (1): The date for Physemothrips should be corrected from 1963a to 1962b, and the reference Stannard (1963a) emended accordingly in the references as per the page "Pacific Insects Issues Online" on the Bishop Museum's website.[4]

Page 19 (2): To match the reference, the authorship for Quirosiella should be spelled "Retana-Salazar" instead of "Retana". It was not possible to verify this by consulting Retana-Salazar (2000), but that spelling is also used by Goldarazena et al. (2008), amongst others.

Page 25: Per the entry for Cricothrips above, the date for Leptogastrothrips should be corrected from 1913 to 1912c.

Page 29: It is quite likely that the date for Adurothrips should be 1994 instead of 1995. Nomenclator Zoologicus gives the date of issue as "December 20, 1994", and 1994 is also the date given on the Schweizerbart website. The original publication could not be checked for confirmation.

Page 31: The author for Austrothrips should be spelled "Brèthes"

Page 32: Issue 5-6 of Marcellia volume 5 is dated late January 1909. The date for Cecidothrips should be corrected from 1908 to 1909 and the reference emended:

Page 35: The authorship for Hexagoniothrips should be emended from "Johansen & Oca" to "Johansen & Montes de Oca".

Page 40 (1): Amongst the authors of Mirothrips, the name "Prezotto" should be spelled "Prezoto".

Page 40 (3): Correct the author for Moultonides from "Kevin" to "Kevan" and emend the corresponding Kevan (1963) reference.

Page 40: The correct date and cross-reference for Moultonia is 1929f (not 1928).

Page 44: Per the entry for Cricothrips above, the date for Stephanothrips should be corrected from 1913 to 1912c.

Page 46: The correct place and year of publication for Xylaplothrips is not Priesner (1928), but (per ICZN art. 12.2.5) Priesner (1925d).

Misspelled entriesEdit

Page 5: The entry for Pseudoaeolothrips should be spelled Pseudaeolothrips

Page 8: The entry for Dendrothrips (Vicinichaetella) should probably be corrected to Vicinechaetella. It was not possible to consult the original publication to fully verify this.

Page 11: The entry for Piliothrips should be corrected to Papiliothrips. Contrary to the statement in Mound & Tree (2009:11), the spelling "Piliothrips" is not an incorrect original spelling used by Bhatti (2006), and does not appear in that work.

Page 26: The entry for Elaphrothrips (Elaphroxothrips) should be spelled Elaphoxothrips.

Page 36: The entry for Pseudosymphyothrips should probably be corrected to Pseudosymphothrips. It was not possible to consult the original publication to fully verify this.

Page 43: The entry for Hennigthrips should be spelled Hennigithrips

Other nonbibliographic emendationsEdit

Page 14: In the entry for Moundiella, add "Junior homonym of Moundiella David, 1974 [Homoptera, Aleyrodidae]."

Page 17: In the entry for Stylothrips, add "Junior homonym of Stylothrips Bondar, 1924."

Page 22: In the entry for Athrips, add "Junior homonym of Athrips Billberg, 1820 [Lepidoptera, Gelechiidae]."

Page 25 (1): The first author to synonymize Leurothrips and Compsothrips was not Priesner (1928a) but Bagnall (1909a).

Page 25 (2): Macrothrips Buffa should also have a note added: "Junior homonym of Macrothrips Bagnall, 1908". Its cross-references should also be corrected from 1908 to 1908a.

Page 28: The entry for Meiothrips (Aculeathrips) should be split to account for its replaced homonym:

Meiothrips (Telothrips) Kudo & Ananthakrishnan, 1974. Type species Meiothrips (Telothrips) nepalensis Kudo & Ananthakrishnan, by monotypy.
Meiothrips (Aculeathrips) Kudo, 1975. Replacement name for Meiothrips (Telothrips) Kudo & Ananthakrishnan, 1974, not Telothrips Priesner, 1929a. Synonymy by Palmer & Mound (1978).

Page 30: Apelaunothrips is also a replacement name whose entry requires splitting:

Apelaunothrips Karny, 1925b [40 species]. Replacement name for Ophidothrips Karny, 1925c, not Ophidothrips Schmutz, 1909.
Ophidothrips Karny, 1925c. Type species Ophidothrips medioflavus Karny, 1925c, by monotypy.

Page 38: Under Aguilaria, replace the mention "nomen nudum" with "Incorrect original spelling for Dioclesianothrip".

Page 41: In the entry for Phlaeothrips (Odontinothrips), the second parenthesis is missing.

Cross-references to be correctedEdit

Throughout: All cross references to Priesner (1927a) should be corrected to 1927.

Page 5 (1): The cross-references for Audiothrips should be corrected to 1930a.

Page 5 (2): The cross-references for Corynothripoides should be corrected to 1926a.

Page 6 (1): The cross-reference for both Fauriella and Opisthothrips should be corrected from 1937a to 1937c.

Page 6 (2): The correct cross-reference for Stomatothrips is Hood, 1912b.

Page 8: The cross-reference for Stenurothrips should be corrected from 1914d to 1914e.

Page 10: The cross-reference for Dinurothrips should be corrected from 1913b to 1913d.

Page 11 (1): The cross-references for Stosicthrips should be corrected from 2009 to 2009b.

Page 11 (2): The cross-references for Sensothrips should be corrected from 1999d to 1999b.

Page 12 (1): The cross-references for Amorphothrips should be corrected from 1924 to 1924d.

Page 12 (2): The cross-references for Amphorothrips should be corrected from 1939c to 1939d.

Page 13: The cross-references for Croceithrips should be corrected from 1995a to 1995b.

Page 14 (1): The cross-references for Craspedothrips should be corrected from 1966 to 1966b.

Page 14 (2): The cross-references for Craspedothrips (Antenothrips) should be corrected from 1995 to 1995b.

Page 19: The cross-references for Procerothrips should be corrected from 1924 to 1924d.

Page 22: The cross-references for Bacillothrips should be corrected from 1908 to 1908b.

Page 27: The cross-references for Siphonothrips should be corrected from 1908 to 1908b.

Page 28: The cross-references for Polytrichothrips should be corrected from 1939 to 1939b.

Page 30 (1): The cross-references for Amphibolothrips should be corrected from 1909 to 1909b.

Page 30 (2): The cross-references for Ampothrips should be corrected from 1939 to 1939d.

Page 33: The cross-references for Ecacanthothrips should be corrected from 1909 to 1909a.

Page 34: The cross-references for Heterothrips should be corrected from 1908 to 1908c.

Page 37 (1): The cross-references for Odontoplothrips should be corrected from 1928d to 1928e.

Page 37 (2): The cross-references for Jacobothrips should be corrected from 1966 to 1966a.

Page 39: The cross-references for Rhynchothrips should be corrected from 1912c to 1912e.

Page 41: The cross-references for Philothrips should be corrected from 1939 to 1939d.

Page 43: The cross-references for Rhaptothrips should be corrected from 1909 to 1909b.

Page 44: The cross-references for Smicrotrhips should be corrected from 1952a to 1952.

Missing referencesEdit

  • Bagnall, R.S. (1924c). Von Schlechtendal's work on fossil thysanoptera in the light of recent knowledge. Annals and Magazine of Natural History (9). 14(79): 156-162.
  • Bagnall, R.S. (1924d). Fossil Thysanoptera III. Terebrantia Pt. 3. Entomologist's monthly Magazine. 60: 251-252.
  • Buffa, P. (1908a) Esame di una piccola raccolta di Tisanotteri esistente nel Museo zoologico della R. Università di Napoli. Annuario del Museo Zoologico della Università di Napoli (N.S.), 2(23): 1-8.
  • Buffa, P. (1908c) Tisanotteri nuovo, nota preliminare. Redia, 5(1): 123-125.
  • Buffa, P. (1909) Contribuzione alla conoscenza dei Tisanotteri (due nuovi generi di Tubuliferi). Bollettino del Laboratorio di zoologia generale e agraria della R. Scuola superiore d'agricoltura in Portici, 3: 193-199.
  • Burmeister, H. (1838) Handbuch der Entomologie. Zweitter Bande: Besondere Entomologie. Zweite Abtheilung: Kaukerfe. Gymnognatha. Erste Hälfte: vulgo Orthoptera. Theod. Chr. Friedr. Enslin, Berlin. viii+397-756 pp.
  • Crawford, D.L. (1909a) Some new Thysanoptera from southern California, I. Pomona College Journal of Entomology. 1: 100-108.
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  • Hood, J. D. (1937c). New genera and species of Thysanoptera from South Africa. Annals and Magazine of Natural History (10). 19(109): 97-113.
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Other bibliographical emendationsEdit

Full updated references are only given where more than one element requires emendation.

Ananthakrishnan (1967) to Annales de la société entomologique de France (N.S.)

Ananthakrishnan (1969a) to 2 [1968], 201–204.1

Bagnall (1909a) to Annales de la Société entomologique de Belgique

Bhatti (1979d) to Nouvelle Revue d’Entomologie

There are three references labelled Bhatti (1995a), which are referenced with correctly labelled different cross-references, they require emending as follow:

Bhatti, J.S. (1995a) Moundothrips, a new genus of Thripidae from Java, Indonesia (Insecta: Terebrantia). Zoology, 5 (1), 65–72.
Bhatti, J.S. (1995b) Further studies on Taeniothrips sensu lato (Insecta: Terebrantia: Thripidae). Zoology, 5 (1), 73–94.
Bhatti, J.S. (1995c) Studies on some Phlaeothripidae (Insecta:Tubulifera). Zoology, 5 (1), 97–110.

Bournier (1962) to:

Bournier, A. (1962). Thysanoptères de France II. Descriptions sommaires de quelques espèces nouvelles du Languedoc méditerranéen. Bulletin de la Société entomologique de France. 67(1): 41-43.

Bournier & Bournier (1986) to Annales de la Société d’Horticulture et d’Histoire Naturelle de l’Hérault, 126(3): 42-45

Brethes (1915) to

Brèthes, J. (1915) Descripcion de un Genero nuevo y una nueva especie de Thysanoptero de la Republica Argentina.

Anales del Museo Nacional de Historia Natural de Buenos Aires, 27, 89–92.

Buffa (1907) to Atti della Società Toscana di Scienze Naturali. Processi Verbali.

Buffa (1908) to 1908b

Crawford (1909) to:

Crawford, D.L. (1909b). Some Thysanoptera of Mexico and the South I. Pomona College Journal of Entomology. 1: 109-119.

Franklin (1907) to 18: 247-250.

Hood (1913a) by deleting the DOI

John (1921) to 1922.

John (1929) to Bulletin et Annales de la Société Entomologique de Belgique

Karny (1908) to "Die zoologische Reise des naturwissenschaftlichen Vereines nach Dalmatien im April 1906. B. Spezieller Teil. Bearbeitung des gesammelten Materials. 6. Orthoptera und Blattaeformia"

Karny (1912b) to 11: 115-169

Karny (1919) to Neue Beiträge zur systematischen Insektenkunde

Kieffer (1908) to 1909

Maltbaek (1928) is a puzzling rference. the name Priesneria doe snot appear in the article cited in the Catalogue, the following reference is tentatively reconstructed from the references in that article and the citation given in zur Strassen (1996)

Maltbaek, J. (1928) Frynsevinger (Thysanoptera). Et Bidrag til Kundskaben om Haderslev-Egnens Fauna. Indbydelsessk. Afgangs. Aarspr. Haderslev Katedralskoles Aarsskrif VI: 1927:I-II.

Marchal (1910) to 2(1): 17-20.

Morison (1949) to:

Morison, G.D. (1949) Thysanoptera of the London Area. Part III. Key to British genera and species of Thysanoptera contd.). The London Naturalist, 28 (suppl.): 77–131.

Moulton (1930b) to Revista Chilena de Historia Naturale. 34: 272–275.

Moulton (1949) to (12) 2, 481–498

Mound & Palmer (1981) to Entomologica Scandinavica Supplement. 15: 153–170.

Okajima (1989) to 1990

In Oustalet (1873), the "M" is the French abbreviation for "Monsieur", the author is Émile Oustalet, otehrwise better known as an ornithologist and mammalogist.

Oustalet, É. (1873) Sur quelques espèces fossiles de l’ordre des Thysanoptères. Bulletin de la Société Philomathique de Paris (6), 10: 20–27.

Priesner (1920a) to Sitzungsberichte der Mathematisch-Naturwissenschaftliche Klasse der Kaiserlichen Akademie der Wissenschaften

Priesner (1921c) to Deutsche entomologische Zeitschrift and

Priesner (1922b) to Sitzungsberichte der Mathematisch-Naturwissenschaftliche Klasse der Kaiserlichen Akademie der Wissenschaften

Priesner (1924b) by deleting the DOIPriesner (1925b) to to

Priesner (1927) to 1927a

Priesner (1929c) from “Metawiensia” to “Mentawiensia”

Priesner (1932e) to Revue de zoologie et de botanique africaines, 22, 192–221; 320–344.

Priesner (1940a) 10 71: 403–409.

Priesner (1953) by deleting the DOI

Priesner (1965) to Publications de l’Institut du Désert d’Égypte

Ramakrishna & Margabandhu (1931) to 1029-1040

Retana-Salazar (2001) is erroneously cited in either the Catalogue or the website of the National Library of Costa Ricat. Assuming the latter is correct, this should be emended to 53: 51-621

Reuter (1904) to Meddelanden af Societas pro Fauna et Flora Fennica

Schille (1911) to Sprawozdanie Komisyi Fizyograficznej, Sekcyę Zoologiczną

Schliephake (1999) to

Schmutz (1913) to Sitzungsberichte der Mathematisch-Naturwissenschaftliche Klasse der Kaiserlichen Akademie der Wissenschaften

Stannard (1952a) to 65: 127-130

Trybom (1913) by deleting the DOI

Vuillet (1914a) to 1914(6): 189-190 or 19(6): 189-190 depending on one's choice, but "6" alone is not an appropriate volume number here.

Williams (1913) to Journal of Economic Biology




I wish to thank Lawrence Mound for early discussion of some issues detailed in this article and for giving his benediction for its publication. Dr. Mound did not review this article prior to publication. I also want to acknowledge Doug Yanega for helping with some of the thornier nomenclatural issues.


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  1. Ich habe deshalb den Serville sehen Namen in diesem Sinne gebraucht, um nicht einen neuen Namenin die Literatur einführen zu müssen.
  2. See Skelley (2020)'s discussion of Tritomapara for a example of the consequences that using Art. 60.3 to replace a misinterpreted genus name can have.
  3. While the incorrect subsequent spelling is in prevalent usage, it is attributed to Priesner, and is technically not protected under ICZN article 33.3.1 even though Priesner was until now universally assumed to the be original author.