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- Wang, H.S., Holloway, J.D., Wahlberg, N., Wang, M. & Nylin, S. 2019. Molecular phylogenetic and morphological studies on the systematic position of Heracula discivitta reveal a new subfamily of Pseudobistonidae (Lepidoptera: Geometroidea). Systematic Entomology 44(1): 211–225. DOI: 10.1111/syen.12326
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