Template:Verdcourt & Bridson, 1991

• Verdcourt, B. & Bridson, D. 1991. Rubiaceae (Part 3). In Polhill, R.M. (ed.), Flora of Tropical East Africa. A.A.Balkema/Rotterdam/Brookfield, pp. 749–956. ISBN 90 6191 357 8 Google Books.  
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