Template:Toussaint et al., 2016a
- Toussaint, E.F.A., Balke, M., García, M. & Short, A.E.Z. 2016. Molecular Systematics of the Neotropical Diving Beetle Genus Rugosus García, 2001 (Coleoptera: Dytiscidae: Copelatinae). The Coleopterists Bulletin 70(1): 53–58. DOI: 10.1649/072.070.0105 . JSTOR .