Template:Testo, 2018

• Testo, W.L. 2018. Novelties in Costa Rican Pityrogramma (Pteridaceae): A New Species and a New Hybrid from the Osa Peninsula. American Fern Journal 108(1): 27–33, DOI: 10.1640/0002-8444-108.1.27 ResearchGate  
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