Sasaji, H. & Kishimoto, O. 1996. Insect fauna of Nakaikemi Marsh, Fukui Prefecture, Central Japan and its natural environmental conservation, with description of a new species and its life history (Insecta: Coleoptera: Coccinellidae). The Memoirs of the Snowdraft Laboratory (Fukui University) 3: 15-36.
Kovář, I. 2007: Family Coccinellidae Latreille, 1807. Pp. 568–631 in: Löbl, I. & Smetana, A. (eds.) Catalogue of Palaearctic Coleoptera. Volume 4: Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Strenstrup. Full text PDFReference page.
