Template:Pool, 2005
- Pool, A. 2005. Urticaceae. Pp. 391–402 in Steyermark, J.A. (†), Berry, P.E., Yatskievych, K. & Holst, B.K. (eds.), Flora of the Venezuelan Guayana, Vol. 9, Rutaceae–Zygophyllaceae. Missouri Botanical Garden Press, St. Louis. ISBN 1-930723-47-4.