Template:Maddison & Toledano, 2012
- Maddison, D.R.; Toledano, L. 2012: A new species of Bembidion (Ecuadion) from Ecuador (Coleoptera, Carabidae, Bembidiini), with a key to members of the georgeballi species group. ZooKeys 249: 51–60. DOI: 10.3897/zookeys.249.4149