Template:Lees et al., 2003a
- Lees, D.C., Kremen, C. & Raharitsimba, H. 2003. Classification, diversity, and endemism of the butterflies (Papilionoidea and Hesperioidea): a revised species checklist. In: Goodman, S.M., & Benstead, J.P. [Eds]. The Natural History of Madagascar. University of Chicago Press, 1709 pp. Chapter pagination: 762–793.