Template:Josso, 2011
- Josso, J.-F. 2011: Description of four new species of Scarabaeidae from Zambia belonging to the genus Catharsius Hope, 1837, Copris Muller, 1764, Pleuronitis Lansberge, 1875 and Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Coprini, Onitini, Onthophagini). Catharsius, (3): 1–8. [not seen]