Template:Howden, 1982
- Howden, H.F. 1982: Larval and adult characters of Frickius Germain, its relationship to the Geotrupini, and a phylogeny of some major taxa in the Scarabaeoidea (Insecta: Coleoptera). Canadian journal of zoology 60(11): 2713–2724. DOI: 10.1139/z82-347