Template:Harley, 1999
- Harley, R.M. 1999. Lamiaceae. Pp. 678–700 in Steyermark, J.A. (†), Berry, P.E., Yatskievych, K. & Holst, B.K. (eds.), Flora of the Venezuelan Guayana, Vol. 5, Eriocaulaceae–Lentibulariaceae. Missouri Botanical Garden Press, St. Louis. ISBN 0-915279-71-1.