Date of publication: 29 April 2009 (online); June 2009 (issue date); 11 June 2009 (issue online)
Note: In the early (pre-print?) version of the article published electronically, "Unpublished for the purposes of zoological nomenclature (Art. 8.2, ICZN)" appears in the footer of the first page; this statement is not in the final version.