Template:Frolov, 2012
- Frolov, A.V. 2012: Diagnosis, classification, and phylogenetic relationships of the orphnine scarab beetles (Coleoptera, Scarabaeidae: Orphninae). Entomologicheskoe obozrenie, 91(2): 332–351. [in Russian, English translation in Entomological review, 92(7): 782–797. (2012) DOI: 10.1134/S0013873812070056]