Template:Aymard & Cuello, 2004
- Aymard C., G.A. & Cuello A., N.L. 2004. Rosaceae. Pp. 490–496 in Steyermark, J.A. (†), Berry, P.E., Yatskievych, K. & Holst, B.K. (eds.), Flora of the Venezuelan Guayana, Vol. 8, Poaceae–Rubiaceae. Missouri Botanical Garden Press, St. Louis. ISBN 1-930723-36-9.