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NOTE: According to Maddison et al. (2008: 16), Carabidae is probably paraphyletic with respect to the other families of Caraboidea as listed here (i.e., Cicindelidae, Rhysodidae, and Trachypachidae), so a complete phylogenetic reclassification of the superfamily is needed.


  • Maddison, D.R.; Moore, W.; Baker, M.D.; Ellis, T.M.; Ober, K.A.; Cannone, J.J.; Gutell, R.R. 2008: Monophyly of terrestrial adephagan beetles as indicated by three nuclear genes (Coleoptera: Carabidae and Trachypachidae). Zoologica scripta, doi: 10.1111/j.1463-6409.2008.00359.x
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