Mark Stephen Harvey

Mark Stephen Harvey (born 1958), Australian arachnologist.

• Department of Terrestrial Zoology Western Australian Museum, Locked Bag 49 Welshpool DC, Western Australia 6986, Australia.
• Division of Invertebrate Zoology American Museum of Natural History; California Academy of Sciences, San Francisco, USA.
• School of Animal Biology University of Western Australia, Crawley Western Australia 6009, Australia.
• School of Natural Sciences Edith Cowan University, Joondalup Western Australia 6027, Australia.
• E-mail: mark.harvey@museum.wa.gov.au

Taxon names authored

(List may be incomplete)

• 289 taxon names authored by Mark Stephen Harvey

Publications

(List may be incomplete)

1985

• Harvey, M.S. & Mahnert, V. 1985. Olpium L. Koch, 1873 (Arachnida, Pseudoscorpionida, Olpiidae): proposed designation of type species and related problems. Bulletin of Zoological Nomenclature 42: 85–88. Reference page. 

1989

• Harvey, M.S. 1989a. A new species of Feaella Ellingsen from north-western Australia (Pseudoscorpionida: Feaellidae). Bulletin of the British Arachnological Society 8: 41–44. Reference page. 

1992

• Harvey, M.S. 1992. The Schizomida (Chelicerata) of Australia. Invertebrate taxonomy 6(1): 77–129. DOI: 10.1071/IT9920077 Reference page. 
• Harvey, M.S. 1992. A new species of Symphytognatha Hickman (Araneae: Symphytognathidae) from Western Australia. Records of the Western Australian Museum 15: 685–689. Reference page. 

2001

• Harvey, M.S. 2001. Notes on the spider genus Symphytognatha (Araneae: Symphytognathidae) in Western Australia. Records of the Western Australian Museum 20: 345–347. Reference page. 

2006

• Harvey, M.S. & Mahnert, V. 2006. The systematic position of the Amazonian species of Albiorix (Pseudoscorpiones, Ideoroncidae). Journal of Arachnology 34: 227–230. Reference page. 
• Šťáhlavský, F., Král, J., Harvey, M.S. & Haddad, C.R. 2006. A karyotype study on the pseudoscorpion families Geogarypidae, Garypinidae and Olpiidae (Arachnida: Pseudoscorpiones). European Journal of Entomology 103: 277–289. Reference page. 

2007

Harvey, Barba, Muchmore & Pérez 2007. Pseudalbiorix, a new genus of Ideoroncidae (Pseudoscorpiones, Neobisioidea) from central America. Journal of Arachnology 34: 610–626. Reference page. 

2009

• Edward, K.L. & Harvey, M.S. 2009. Australian goblin spiders of the genus Ischnothyreus (Araneae, Oonopidae). Records of the Western Australian Museum 25(3): 287–293 full article (PDF). Reference page. 

2010

• Burger, M., Harvey, M.S. & Stevens, N.B. 2010. A new species of blind subterranean Tetrablemma (Araneae: Tetrablemmidae) from Australia. The Journal of Arachnology 38: 146–149. Reference page. 
• Framenau, V.W.; Scharff, N.; Harvey, M.S. 2010: Systematics of the Australian orb-weaving spider genus Demadiana with comments on the generic classification of the Arkyinae (Araneae: Araneidae). Invertebrate systematics, 24(2): 139–171. DOI: 10.1071/IS10005Reference page. 
• Harvey, M.S. 2010. Grymeus, a new genus of pouched oonopid spider from Australia (Chelicerata: Araneae). Memoirs of the Museum of Victoria 48(2): 123-130. full article (PDF). Reference page. 

2011

• Crews, S.C. & Harvey, M.S. 2011. The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region. ZooKeys 99: 1–103. DOI: 10.3897/zookeys.99.723. Reference page. 
• Harvey, M.S. 2011. Order Pseudoscorpiones de Geer, 1778. Pp 119–120 In Zhang, Z.-Q. (ed.) 2011. Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 1–237. Open access. Reference page. . Full article (PDF) Reference page. 
• Rix, M.G. & Harvey, M.S. 2011. Australian assassins, part I: a review of the assassin spiders (Araneae: Archaeidae) of mid-eastern Australia. ZooKeys 123: 1-100. DOI: 10.3897/zookeys.123.1448 Reference page. 

2012

• Baehr, B.C., Harvey, M.S., Burger, M. & Thoma, M. 2012. The new Australasian goblin spider genus Prethopalpus (Araneae, Oonopidae). Bulletin of the American Museum of Natural History, (369). 113pp. DOI: 10.1206/763.1 . hdl: 2246/6185 . Reference page. 
• Harvey, F.S.B., Framenau, V.W., Wojcieszek, J.M., Rix, M.G. & Harvey, M.S. 2012. Molecular and morphological characterisation of new species in the trapdoor spider genus Aname (Araneae: Mygalomorphae: Nemesiidae) from the Pilbara bioregion of Western Australia. Zootaxa 3383: 15–38. Preview PDF. Reference page. 
• Šťáhlavský, F., Boyer, S.L., Harvey, M.S. & Giribet, G. 2012. First cytogenetic study of a member of the harvestman family Pettalidae (Opiliones: Cyphophthalmi). Australian Journal of Entomology, DOI: 10.1111/j.1440-6055.2012.00870.x Reference page. 
• Šťáhlavský, F., Král, J., Harvey, M.S. & Haddad, C.R. 2012a. The first cytogenetic characterisation of atemnids: pseudoscorpions with the highest chromosome numbers (Arachnida: Pseudoscorpiones). Cytogenetic and Genome Research 137: 22–30. Reference page. 

2013

• Baehr, B.C., Harvey, M.S., Smith, H.M. & Ott, R. 2013. The goblin spider genus Opopaea in Australia and the Pacific islands (Araneae: Oonopidae). Memoirs of the Queensland Museum, Nature 58: 107-338. Reference page. 
• Car, C.A., Short, M., Huynh, C. & Harvey, M.S. 2013. The millipedes of Barrow Island, Western Australia (Diplopoda). Records of the Western Australian Museum, Supplement 83: 209–219. DOI: 10.18195/issn.0313-122x.83.2013.209-219 . Reference page. 
• Gunawardene, N.R., Majer, J.D., Taylor, C.K. & Harvey, M.S. (eds.) 2013. The terrestrial invertebrate fauna of Barrow Island, Western Australia. Records of the Western Australian Museum, Supplement 83. Reference page. 
• Harms, D. & Harvey, M.S. 2013. Review of the cave-dwelling species of Pseudotyrannochthonius Beier (Arachnida: Pseudoscorpiones: Pseudotyrannochthoniidae) from mainland Australia, with description of two troglobitic species. Australian journal of entomology 52(2): 129–143. DOI: 10.1111/aen.12009 Reference page. 
• Harvey, M.S. & Muchmore, W.B. 2013: The systematics of the pseudoscorpion family Ideoroncidae (Pseudoscorpiones: Neobisioidea) in the New World. Journal of arachnology 41(3): 229–290. DOI: 10.1636/K13-42.1 Reference page. 
• Harvey, M.S. 2013. Order Pseudoscorpiones. In: Zhang, Z.-Q. (ed.) 2013. Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness (Addenda 2013). Zootaxa 3703(1): 34–35. DOI: 10.11646/zootaxa.3703.1.8 . Reference page. 
• Humphreys, G., Alexander, J., Harvey, M.S. & Humphreys, W.F. 2013. The subterranean fauna of Barrow Island, north-western Australia: 10 years on. Records of the Western Australian Museum, Supplement 83: 145–158. DOI: 10.18195/issn.0313-122x.83.2013.145-158 . Reference page. 
• Majer, J.D., Gunawardene, N.R., Taylor, C.K. & Harvey, M.S. 2013. A last word. Records of the Western Australian Museum, Supplement 83: 405–406. DOI: 10.18195/issn.0313-122x.83.2013.405-406 . Reference page. 

2014

• Edward, K.L. & Harvey, M.S. 2014. Australian goblin spiders of the genus Ischnothyreus (Araneae, Oonopidae). Bulletin of the American Museum of Natural History 389: 1–144 DOI: 10.1206/865.1 Reference page. 
• Giribet, G., McIntyre, E., Christian, E., Espinasa, L., Ferreira, R., Francke, Ó.F., Harvey, M.S., Isaia, M., Kováč, Ľ., McCutchen, L., Souza, M.F.V.R. & Zagmajster, M. 2014. The first phylogenetic analysis of Palpigradi (Arachnida) – the most enigmatic arthropod order. Invertebrate Systematics 28(4): 350–360. DOI: 10.1071/IS13057 Reference page. 
• Miglio, L.T., Harms, D., Framenau, V.W. & Harvey, M.S. 2014. Four new mouse spider species (Araneae, Mygalomorphae, Actinopodidae, Missulena) from Western Australia. ZooKeys 410: 121–148. DOI: 10.3897/zookeys.410.7156 . Reference page. 

2015

• Ott, R., von Beeren, C., Hashim, R., Witte, V. & Harvey, M.S. 2015a. Sicariomorpha, a new myrmecophilous goblin spider genus (Araneae, Oonopidae) associated with Asian army ants. American Museum Novitates 3843: 1-14. DOI: 10.1206/3843.1. . Reference page. 

2016

• Giribet, G., Boyer, S.L., Baker, C.M., Fernández, R., Sharma, P., de Bivort, B.L., Daniels, S.R.., Harvey, M.S. & Griswold, C.E. 2016. A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling. Zoological Journal of the Linnean Society 178: 523–545. Reference page. 
• Harvey, M.S., Huey, J.A., Hillyer, M.J., McIntyre, E. & Giribet, G. 2016a. The first troglobitic species of Gymnobisiidae (Pseudoscorpiones, Neobisioidea), from Table Mountain (Western Cape Province, South Africa) and its phylogenetic position. Invertebrate Systematics 30: 75–85. Reference page. 
• Harvey, M.S., Abrams, K.M., Beavis, A.S., Hillyer, M.J. & Huey, J.A. 2016. Pseudoscorpions of the family Feaellidae (Pseudoscorpiones : Feaelloidea) from the Pilbara region of Western Australia show extreme short-range endemism. Invertebrate Systematics 30: 491–508. Reference page. 

2017

• Car, C.A. & Harvey, M.S. 2017. New species of Boreohesperus (Diplopoda, Polydesmida, Paradoxosomatidae) from north-western Australia. European Journal of Taxonomy 320: 1–11. full article (PDF). Reference page. 
• Castalanelli, M.A., Huey, J.A., Hillyer, M.J. & Harvey, M.S. 2017. Molecular and morphological evidence for a new genus of small trapdoor spiders from arid Western Australia (Araneae: Mygalomorphae: Nemesiidae: Anaminae). Invertebrate Systematics 31(4): 492-505. DOI: 10.1071/IS16061. Reference page. 
• Gao, Z.Z., Zhang, F. & Harvey, M.S. 2017. A modified definition of the genus Haplochernes (Pseudoscorpiones: Chernetidae), with a new species from Hainan Island. Journal of Arachnology 45(1): 112-122. Reference page. 
• Rix, M.G., Raven, R.J., Main, B.Y., Harrison, S.E., Austin, A.D., Cooper, S.J.B. & Harvey, M.S. 2017. The Australasian spiny trapdoor spiders of the family Idiopidae (Mygalomorphae : Arbanitinae): a relimitation and revision at the generic level. Invertebrate Systematics 31(5): 566-634. DOI: 10.1071/IS16065. Reference page. 
• Smith, H.M., Harvey, M.S., Agnarsson, I. & Anderson, G.J. 2017. A Revision and Cladistic Analysis of the Genus Corasoides Butler (Araneae: Desidae) with Descriptions of Nine New Species. Records of the Australian Museum 69(10: 15–64. DOI: 10.3853/j.2201-4349.69.2017.1672. . Full article (PDF). Reference page. 

2018

• Harrison, S.E., Rix, M.G., Harvey, M.S. & Austin, A.D. 2018. Systematics of the Australian spiny trapdoor spiders of the genus Blakistonia Hogg (Araneae: Idiopidae). Zootaxa 4518(1): 001–076. DOI: 10.11646/zootaxa.4518.1.1. Reference page. 
• Harvey, M.S., Hillyer, M.J., Main, B.Y., Moulds, T.A., Raven, R.J., Rix, M.G., Vink, C.J. & Huey, J.A. 2018. Phylogenetic relationships of the Australasian open-holed trapdoor spiders (Araneae: Mygalomorphae: Nemesiidae: Anaminae): multi-locus molecular analyses resolve the generic classification of a highly diverse fauna. Zoological Journal of the Linnean Society 184(2): 407-452. DOI: 10.1093/zoolinnean/zlx111. Reference page. 
• Novák, J. & Harvey, M.S. 2018. New species and records of the pseudoscorpion genus Geogarypus (Pseudoscorpiones: Geogarypidae) from India, Sri Lanka and New Guinea. Zootaxa 4394(3): 417–427. DOI: 10.11646/zootaxa.4394.3.7 Reference page. 
• Rix, M.G., Huey, J.A., Cooper, S.J.B., Austin, A.D. & Harvey, M.S. 2018. Conservation systematics of the shield-backed trapdoor spiders of the nigrum-group (Mygalomorphae, Idiopidae, Idiosoma): integrative taxonomy reveals a diverse and threatened fauna from south-western Australia. ZooKeys 756: 1–121. DOI: 10.3897/zookeys.756.24397. Reference page. 
• Rix, M.G., Raven, R.J., Austin, A.D., Cooper, S.J.B. & Harvey, M.S. 2018. Systematics of the spiny trapdoor spider genus Bungulla (Mygalomorphae: Idiopidae): revealing a remarkable radiation of mygalomorph spiders from the Western Australian arid zone. The Journal of Arachnology 46(2): 249-344. DOI: 10.1636/JoA-S-17-057.1. . Reference page. 
• Rix, M.G., Main, B.Y., Raven, R.J. & Harvey, M.S. 2018. Systematics of the spiny trapdoor spiders of the genus Eucanippe (Mygalomorphae: Idiopidae: Aganippini) from south-western Australia: documenting a poorly-known lineage from Australia's biodiversity hotspot. The Journal of Arachnology 46(1): 133-154 & Appendix "Atlas of male morphology spiny trapdoor spiders (genus Eucanippe)". DOI: 10.1636/JoA-S-17-030.1. . Reference page. 
• Rix, M.G., Raven, R.J. & Harvey, M.S. 2018. Systematics of the giant spiny trapdoor spiders of the genus Gaius Rainbow (Mygalomorphae: Idiopidae: Aganippini): documenting an iconic lineage of the Western Australian inland arid zone. The Journal of Arachnology 46(3): 438-472. DOI: 10.1636/JoA-S-17-079.1. . Reference page. 

2019

• Car, C.A., Harvey, M.S., Hillyer, M.J. & Huey, J.A. 2019. The millipede genus Antichiropus (Diplopoda: Polydesmida: Paradoxosomatidae), part 3: species of the Pilbara bioregion of Western Australia. Zootaxa 4617(1): 1–71. DOI: 10.11646/zootaxa.4617.1.1 Reference page. 
• Huey, J.A., Rix, M.G., Wilson, J.D., Hillyer, M.J. & Harvey, M.S. 2019. Open-holed trapdoor spiders of the genus Teyl (Mygalomorphae: Nemesiidae: Anamini) from Western Australia’s Pilbara bioregion: a new species and expanded phylogenetic assessment. Zootaxa 4674(3): 349–362. DOI: 10.11646/zootaxa.4674.3.3. Reference page. 
• Kropf, C., Blick, T., Brescovit, A.D., Chatzaki, M., Dupérré, N., Gloor, D., Haddad, C.R., Harvey, M.S., Jäger, P., Marusik, Y.M., Ono, H., Rheims, C.A. & Nentwig, W. 2019. How not to delimit taxa: a critique on a recently proposed “pragmatic classification” of jumping spiders (Arthropoda: Arachnida: Araneae: Salticidae). Zootaxa 4545(3): 444–446. DOI: 10.11646/zootaxa.4545.3.10. . Reference page. 
• Ott, R., Ubick, D., Bonaldo, A.B., Brescovit, A.D. & Harvey, M.S. 2019. A revision of the new world goblin spider genus Cinetomorpha Simon, 1892 revalidated from Gamasomorpha Karsch, 1881 (Araneae, Oonopidae, Oonopinae). Zootaxa 4641(1): 001–152. DOI: 10.11646/zootaxa.4641.1.1. Reference page. 
• Rix, M.G., Wilson, J.D. & Harvey, M.S. 2019. A revision of the white-headed spiny trapdoor spiders of the genus Euoplos (Mygalomorphae: Idiopidae: Arbanitinae): a remarkable lineage of rare mygalomorph spiders from the south-western Australian biodiversity hotspot. The Journal of Arachnology 47(1): 63-76. Reference page. .
• Rix, M.G., Wilson, J.D., Rix, A.G., Wojcieszek, A.M., Huey, J.A. & Harvey, M.S. 2019. Population demography and biology of a new species of giant spiny trapdoor spider (Araneae: Idiopidae: Euoplos) from inland Queensland: developing a ‘slow science’ study system to address a conservation crisis. Austral Entomology 58(2): 282-297. Reference page. 
• Romero-Ortiz, C. & Harvey, M.S. 2019. The pseudoscorpion genus Verrucachernes (Pseudoscorpiones: Chernetidae) in the Indian region. Zootaxa 4568(2): 337–346. DOI: 10.11646/zootaxa.4568.2.8 Reference page. 

2020

• Castalanelli, M.A., Framenau, V.W., Huey, J.A., Hillyer, M.J. & Harvey, M.S. 2020. New species of the open-holed trapdoor spider genus Aname (Araneae: Mygalomorphae: Anamidae) from arid Western Australia. Journal of Arachnology 48(2): 169-213. DOI: 10.1636/0161-8202-48.2.169. . Reference page. 
• Harvey, M.S., Gruber, K., Hillyer, M.J. & Huey, J.A. 2020. Five new species of the open-holed trapdoor spider genus Aname (Araneae: Mygalomorphae: Anamidae) from Western Australia, with a revised generic placement for Aname armigera. Records of the Western Australian Museum 35: 10-38. DOI: 10.18195/issn.0312-35 3162.35.2020.010-038. Reference page. 
• Rix, M.G., Wilson, J.D. & Harvey, M.S. 2020. First phylogenetic assessment and taxonomic synopsis of the open-holed trapdoor spider genus Namea (Mygalomorphae: Anamidae): a highly diverse mygalomorph lineage from Australia’s tropical eastern rainforests. Invertebrate Systematics 34(7): 679-726. DOI: 10.1071/IS20004. . Reference page. 

2021

• Marek, P.E., Buzatto, B.A., Shear, W.A., Means, J.C., Black, D.G., Harvey, M.S. & Rodriguez, J. 2021. The first true millipede—1306 legs long. Scientific Reports 11(1), #23126: 1–8. DOI: 10.1038/s41598-021-02447-0 Reference page. 

2022

• Harvey, M.S. 2022. Three new species of the pseudoscorpion genus Synsphyronus (Pseudoscorpiones: Garypidae) from semi-arid Western Australia. Australian Journal of Taxonomy 5: 1–15. Reference page. 
• Harvey, M.S. & Harms, D. 2022. The pseudoscorpion genus Centrochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) from central Asia and description of a new species from Nepal. Journal of Arachnology 50: 158–174. Reference page. 
• Harvey, M.S., Wilson, J.D. & Rix, M.G. 2022. Two new species of the open-holed trapdoor spider genus Aname (Araneae: Mygalomorphae: Anamidae) from southern Australia. Australian Journal of Taxonomy 2: 1-8. DOI: 10.54102/ajt.h7iv8. . Reference page. 
• Rix, M.G. & Harvey, M.S. 2022. A new species of the spiny trapdoor spider genus Eucyrtops (Mygalomorphae: Idiopidae) from south-western Australia. Australian Journal of Taxonomy 4: 1-6. DOI: 10.54102/ajt.unvpo. . Reference page. 
• Rix, M.G., Wilmer, J.W. & Harvey, M.S., 2022. Rainforest pelican spiders (Archaeidae: Austrarchaea) of south-eastern Queensland, Australia: two new species and a distributional reassessment of regional endemic clades. Journal of Arachnology 50(2): 231-249. DOI: 10.1636/JoA-S-21-071. . Reference page. 
• Wilson, J.D., Harvey, M.S. & Rix, M.G. 2022. Euoplos eungellaensis (Idiopidae), a new golden trapdoor spider from central-eastern Queensland. Australian Journal of Taxonomy 5: 1-8. DOI: 10.54102/ajt.lmcbc. . Reference page. 
• You, J.Y., Yoo, J.S., Harvey, M.S. & Harms, D. 2022. Some cryptic Korean karst creatures: revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea. Journal of Arachnology 50: 135–157. Reference page. 

2023

• Harvey, M.S., Burger, M.A.A., Abrams, K.M., Finston, T.L., Huey, J.A. & Perina, G. 2023. The systematics of the pseudoscorpion genus Indohya (Pseudoscorpiones: Hyidae) in Australia. Zootaxa 5342(1): 001–119. DOI: 10.11646/zootaxa.5342.1.1. . Reference page. 
• Harvey, M.S., Wilson, J.D. & Rix, M.G. 2023. Three new species of the open-holed trapdoor spider genus Kwonkan (Mygalomorphae: Anamidae) from central Australia. Australian Journal of Taxonomy 21: 1-9. DOI: 10.54102/ajt.l6he8. . Reference page. 
• Harvey, M.S., Wilson, J.D. & Rix, M.G. 2023a. Two new species of the open-holed trapdoor spider genus Proshermacha (Araneae: Mygalomorphae: Anamidae) from southern Western Australia Australian Journal of Taxonomy 43: 1-13. DOI: doi:10.54102/ajt.2k58g. . Reference page. 
• Rix, M.G., Wilson, J.D. & Harvey, M.S. 2023. A revision of the open-holed trapdoor spiders of the Teyl damsonoides-group (Mygalomorphae: Anamidae: Teylinae) from south-western Australia. The Journal of Arachnology 51(3): 287-377. DOI: 10.1636/JoA-S-21-077. . Reference page. 
• Romero-Ortiz, C., Sarmiento, C.E. & Harvey, M.S. 2023. A new genus and five new species of pseudoscorpions (Arachnida, Pseudoscorpiones, Withiidae) from Colombia. Zookeys 1184ː 301–326. DOI: 10.3897/zookeys.1184.106698 Reference page. 
• Wilson, J.D., Rix, M.G. & Harvey, M.S. 2023. Description of five new Aname L. Koch, 1873 (Araneae, Anamidae) species collected on Bush Blitz expeditions. European Journal of Taxonomy 890: 1–22. DOI: 10.5852/ejt.2023.890.2247. . Reference page. 
• Wilson, J.D., Rix, M.G. & Harvey, M.S. 2023a. A new spider species of the genus Proshermacha (Mygalomorphae: Anamidae) from the Coolgardie and Murchison bioregions of Western Australia, collected on a Bush Blitz expedition. Australian Journal of Taxonomy 19: 1-7. DOI: 10.54102/ajt.olxwd. . Reference page. 

2024

• Harvey, M.S. & Steinmann, D.B. 2024. A new troglomorphic species of Larca (Pseudoscorpiones, Larcidae) from Colorado. ZooKeys 1198: 279–294. DOI: 10.3897/zookeys.1198.120353 Reference page.