Dirk Ahrens

Dirk Ahrens, Entomologist, Germany.

• Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 160, 53113 Bonn, Germany.
• E-mail: ahrens.dirk_col@gmx.de

Taxon names authored

(List may be incomplete)

• 523 taxon names authored by Dirk Ahrens

Publications

(List may be incomplete)

1995

• Ahrens, D. 1995. Calloserica poggii n. sp. – eine neue Art aus dem Himalaya (Col., Scarabaeoidea, Sericinae). Entomologische Nachrichten und Berichte 39: 137–138
• Ahrens, D. 1995. Taxonomische Studien an Sericinae. Die Gattungen Deroserica Moser, 1915, Microsericaria Nikolaev, 1979, Pachyderoserica Moser, 1920 und Microserica Brenske, 1894. (Insecta: Coleoptera: Melolonthidae). Entomologische Abhandlungen 57: 37–56

1996

• Ahrens, D. 1996. Revision der Sericini des Himalaya und angrenzender Gebiete. Die Gattungen Lasioserica Brenske, 1896 und Gynaecoserica Brenske, 1896. (Coleoptera, Scarabaeoidea). Coleoptera, Schwanfelder Coleopterologische Mitteilungen 16: 1–48.
• Ahrens, D. & Sabatinelli G. 1996. Revision der gattung Nepaloserica (Coleoptera: Melolonthidae). Fragmenta entomologica 28(2): 209–243.

1998

• Ahrens, D. 1998. Revision der Sericinae des Himalaya-Gebiets. Neue Arten der Microserica marginata (Brsk.) – Gruppe (Insecta: Coleoptera: Melolonthidae). Entomologische Abhandlungen 58: 29–51.
• Ahrens, D. 1998. Autoserica caifensis Brenske, 1897- eine vergessene Art aus dem östlichen mediterranean Raum (Insecta: Coleoptera: Melolonthidae). Reichenbachia 32: 267–270.

1999

• Ahrens, D. 1999. New species and records of Gynaecoserica Brenske, 1896, Calloserica Brenske, 1894, Nepaloserica Frey, 1965 and Lasioserica Brenske, 1896 from the Himalayas and adjacent regions. (Coleoptera: Melolonthidae, Sericini). Entomologische Abhandlungen 58: 181–242. ResearchGate . Reference page. 
• Ahrens, D. 1999a. Eine neue Art der Microserica marginata - Gruppe aus Ost-Nepal. (Coleoptera: Melonthidae: Sericini). Entomologische Abhandlungen 58: 243–246. Reference page. 
• Ahrens, D. 1999b. Revision der Gattung Serica (s.str.) MacLeay des Himalaya-Gebiets (Coleoptera, Melolonthidae). Fragmenta Entomologica 31: 205–332. Reference page. 

2000

• Ahrens, D. 2000. New species of Calloserica Brenske, 1894 from Sikkim and Nepal (Coleoptera, Melolonthidae, Sericini). Zoosymposia 22: 815–824.
• Ahrens, D. 2000. The Fauna of the Arabian Peninsula: Sericinae (Coleoptera, Scarabaeoidea). Fauna of Arabia 18: 177–210.
• Ahrens, D. 2000. Zwei neue Nipponoserica-Arten aus dem Himalaya (Coleoptera: Melolonthidae, Sericini). Entomofauna 21: 25–30.
• Ahrens, D. 2000. Eine neue Serica-Art aus dem Dhaulagiri-Himal (Nepal) (Coleoptera, Melolonthidae). Fragmenta entomologica 32: 287–291.
• Ahrens, D. 2000. Weiteres Material der Gattung Lasioserica Brenske, 1896, mit der Beschreibung acht neuer Arten (Coleoptera: Melolonthidae: Sericini). Acta Musei Moraviae, scientiae biologicae 85: 1–28.
• Ahrens, D. 2000. Synopsis der Gattung Gastroserica Brenske, 1897 des ostasiatischen Festlandes (Coleoptera: Melolonthidae: Sericini). Entomologische Abhandlungen 59: 73–121.
• Ahrens, D. 2000. Neue Erkenntnisse zur Herkunft von Autoserica caifensis Brenske, 1897 (Insecta: Coleoptera: Melolonthidae). Linzer biologische Beiträge 32: 71–74.

2001

• Ahrens, D. 2001. Further records of Microserica Brenske, 1894 from the Himalayas with description of a new species of the subgenus Parvulomaladera Ahrens, 1995. Part 1. (Coleoptera, Melolonthidae, Sericini). Lambillionea 101: 216–224.
• Ahrens, D. 2001. Zwei neue Chrysoserica-Arten aus dem Himalaya. (Coleoptera Melolonthidae). Bollettino della Società Entomologica Italiana 133: 133–147.

2002

• Ahrens, D. 2002. Notes on distribution and synonymy of sericid beetles of Taiwan, with descriptions of new species (Coleoptera, Scarabaeoidea: Melolonthidae). Annales historico-naturales Musei nationalis hungarici 94: 53–91.
• Ahrens, D. 2002. Revision der Arten der Microserica viridicollis Arrow, 1913 – Gruppe (Coleoptera, Melolonthinae, Sericini). Linzer biologische Beiträge 34: 383–412.

2003

• Ahrens, D. 2003. The identity of Maladera infuscata (Moser, 1915), with description of a new species from Taiwan (Coleoptera: Scarabaeidae: Sericini). Zootaxa 392: 1–8. Abstract & excerpt Reference page. 
• Ahrens, D. 2003. Maladera affinis (Blanchard, 1850) comb. n. (Coleoptera, Scarabaeoidea, Sericini), an oriental faunal element in the Malagasy region. Deutsche Entomologische Zeitschrift 50: 133–142. DOI: 10.1002/mmnd.20030500113
• Ahrens, D. 2003. Neue Arten aus der Gruppe der Amiserica insperata (Brenske, 1898) (Coleoptera, Scarabaeoidea, Sericini). Spixiana 26: 65–81.
• Ahrens, D. 2003. Zur Identität der Gattung Neoserica Brenske, 1894, nebst Beschreibung neuer Arten (Coleoptera, Melolonthidae, Sericini). Koleopterologische Rundschau 73: 169–226.

2004

• Ahrens, D. 2004. Two new species of Sericini from Western Nepal (Coleoptera, Scarabaeidae, Sericini): 211–213. In: Hartmann, M. and Baumbach, H. (Eds.): Biodiversität und Naturausstattung im Himalaya. Verein der Freunde und Förderer des Naturkundemuseums Erfurt e.V., Erfurt, 389 pp.
• Ahrens, D. 2004. Monographie der Sericini des Himalaya (Coleoptera, Scarabaeidae). Dissertation.de – Verlag im Internet GmbH, Berlin, 534 pp. Reference page. 
• Ahrens, D. 2004. Revision of the Microserica lineata group (Coleoptera: Scarabaeidae: Sericini). Zootaxa 664: 1–14.
• Ahrens, D. 2004. Revisional notes on Sericinae: The taxonomic status of the “Neotropical” genus Rhynchosymmela Frey, 1974 (Coleoptera, Scarabaeidae). Beiträge zur entomologie 54: 31–35.
• Ahrens, D. 2004. Phylogeny and zoogeography of the Sericini in the Himalayan region. PhD thesis, Freie Universität Berlin; 226 pp. [1]

2005

• Ahrens, D. 2005. A taxonomic review on the Serica (s.str.) MacLeay, 1819 species of Asiatic mainland (Coleoptera, Scarabaeidae, Sericini). Nova Supplemeta Entomologica 18: 1–163.
• Ahrens, D. 2005. The diversification of the endemic Himalayan monsoon-season beetles genus Calloserica inferred from a cladistic analysis (Coleoptera: Scarabaeidae: Sericini). Invertebrate systematics 19: 217–230. DOI: 10.1071/IS04030
• Ahrens, D. 2005. A preliminary cladistic analysis of Nipponoserica, with implications on phylogenetic relationships among sericine chafers (Coleoptera, Scarabaeidae). Systematics and biodiversity 3: 265–279. DOI: 10.1017/S1477200005001751
• Ahrens, D. 2005. New material of the genus Lasioserica Brenske, 1896 with the description of four new species (Coleoptera: Scarabaeidae: Sericini). Linzer biologische Beiträge 37/1: 771–781.
• Ahrens, D. & Pacholátko, P. 2005. Zwei neue Arten aus der Gruppe der Amiserica insperata (Brenske, 1898) aus dem nordöstlichen Indien und Myanmar (Coleoptera, Scarabaeidae, Sericini). Koleopterologische rundschau 75: 311–317.

2006

• Ahrens, D. 2006. Cladistic analysis of Maladera (Omaladera): implications on taxonomy, evolution and biogeography of the Himalayan species (Coleoptera: Scarabaeidae: Sericini). Organisms diversity & evolution 6: 1–16. DOI: 10.1016/j.ode.2005.01.002. . Reference page. 
• Ahrens, D. 2006. Additional data on taxonomy and distribution on Sericini of the Himalayas, with description a further new species of Maladera (Coleoptera, Scarabaeidae). In: Hartmann, M. and Baumbach, H. (Eds.): Biodiversität und Naturausstattung im Himalaya. Verein der Freunde und Förderer des Naturkundemuseums Erfurt e.V., Erfurt: 409–418.
• Ahrens, D. 2006. Revision und phylogenetische Analyse der Gattung Pachyserica Brenske, 1897 (Coleoptera, Melolonthidae, Sericini). Revue suisse de zoologie 113: 487–557.
• Ahrens, D. 2006. The phylogeny of the genus Lasiosericainferred from adult morphology – implications on the evolution of montane fauna of the South Asian orogenic belt (Coleoptera: Scarabaeidae: Sericini). Journal of zoological systematics and evolutionary research 44: 34–53. DOI: 10.1111/j.1439-0469.2005.00340.x
• Ahrens, D. 2006. The phylogeny of Sericini and their position within the Scarabaeidae based on morphological characters (Coleoptera: Scarabaeidae). Systematic entomology 31: 113–144. DOI: 10.1111/j.1365-3113.2005.00307.x

2007

• Ahrens, D. 2007. Type species designations of Afrotropical Ablaberini and Sericini genera (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 1496: 53–62. Abstract & excerpt Reference page. 
• Ahrens, D. 2007. Taxonomic changes and an updated catalogue for the Palaearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 1504: 1–51. Abstract & excerpt Reference page. 
• Hunt, T., Bergsten, J., Levkanicova, Z., Papadopoulou, A., St. John, O., Wild, R., Hammond, P.M., Ahrens, D., Balke, M., Caterino, M.S., Gómez-Zurita, J., Ribera, I., Barraclough, T.G., Bocakova, M., Bocak, L. & Vogler, A.P. 2007. A comprehensive phylogeny of beetles reveals the evolutionary origins of a superradiation. Science 318: 1913–1916. DOI: 10.1126/science.1146954 . Reference page. 

2009

• Ahrens, D. & Fabrizi, S. 2009a. A review on the genus Gynaecoserica Brenske, 1897 (Coleoptera, Scarabaeidae, Sericini). Journal of natural history 43(25-26): 1505–1584. DOI: 10.1080/00222930902968809 Reference page. 
• Ahrens, D. & Fabrizi, S. 2009b. New species of Sericini from the Eastern Himalaya and Tibet (Coleoptera, Scarabaeidae). In: Hartmann, M. & Weipert, M. (eds.): Biodiversität und Naturausstattung im Himalaya III. Verein der Freunde und Förderer des Naturkundemuseums Erfurt e.V., Erfurt. pp. 249–284. Reference page. 

2011

• Ahrens, D. & Fabrizi, S. 2011. New species of Sericini from the Himalaya and adjacent mountains (Coleoptera: Scarabaeidae). Bonn zoological bulletin 60(2): 139–164.

2012

• Bai, M., Ahrens, D., Yang, X-K. & Ren, D. 2012. New fossil evidence of the early diversification of scarabs: Alloioscarabaeus cheni (Coleoptera: Scarabaeoidea) from the Middle Jurassic of Inner Mongolia, China. Insect science 19(2): 159–171. DOI: 10.1111/j.1744-7917.2011.01460.x . Reference page. 
• Ahrens, D. 2012. New species and a morphology-based phylogeny of the genus Nepaloserica (Insecta: Coleoptera: Scarabaeidae). Hartmann & Weipert: Biodiversität und Naturausstattung im Himalaya IV: 305-316 full article (PDF). Reference page. 
• Shrestha, J., Eberle, J. & Ahrens, D. 2012. Further data on the distribution of Himalayan Sericini from the collection of Naturel History Museum Erfurt (Insecta: Coleoptera: Scarabaeidae) with description of a new Xenoserica from West-Nepal. Vernate 31: 379–386.

2013

• Bocak, L., Barton, C., Crampton-Blatt, A., Chesters, D., Ahrens, D. & Vogler, A.P. 2014. Building the Coleoptera tree-of-life for >8000 species: composition of public DNA data and fit with Linnaean classification. Systematic Entomology 39(1): 97–110. DOI: 10.1111/syen.12037 . Reference page. 

2014

• Ahrens, D., Liu, W.G., Fabrizi, S., Bai, M. & Yang, X.K. 2014. A taxonomic review of the Neoserica (sensu lato) septemlamellata group (Coleoptera, Scarabaeidae, Sericini). ZooKeys 402: 67–102. DOI: 10.3897/zookeys.402.7360 Reference page. 
• Ahrens, D., Schwarzer, J. & Vogler, A.P. 2014. The evolution of scarab beetles tracks the sequential rise of angiosperms and mammals. Proceedings of the Royal Society (B), 281(1791) DOI: 10.1098/rspb.2014.1470 Reference page. 
• Ahrens, D. et al. 2014: A taxonomic review of the Neoserica (sensu lato) abnormis group (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 439: 27–82. DOI: 10.3897/zookeys.439.8055 Reference page. 
• Fabrizi, S. & Ahrens, D. 2014. A Monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn zoological bulletin Supplementum 61: 1–124. Reference page. 
• Liu, W.G., Bai, M., Yang, X.K. & Ahrens, D. 2014. Towards an improved knowledge of Sericini of the Tibetan highland: new species and records (Coleoptera, Scarabaeidae). Bonn zoological bulletin 63(2): 157–172.

2015

• Liu, W.G., Bai, M., Yang, X.K. & Ahrens, D. 2015. New species and records of the Neoserica (sensu stricto) group (Coleoptera, Scarabaeidae, Sericini). Journal of natural history 49(39–40): 2379–2395. DOI: 10.1080/00222933.2015.1034208
• Rössner, E. & Ahrens, D. 2015. Addenda zur Revision von Omaloplia Schoenherr, 1817 (Coleoptera: Scarabaeidae, Sericini). Entomologische Zeitschrift 125(3): 161–170.
• Liu, W.G., Bai, M., Eberle, J., Yang, X.K. & Ahrens, D. 2015. A phylogeny of Chinese Sericini using mitochondrial and ribosomal RNA genes (Coleoptera: Scarabaeidae). Organisms diversity & evolution 15: 343–350.
• Sabatinelli, G. & Ahrens, D. (2015) New records and a checklist of Sericini from Pakistan (Coleoptera, Scarabaeoidea, Scarabaeidae). Revista gaditana de Entomología 6(1): 137–150.
• Eberle, J., Walbaum, W., Fabrizi, S. Ahrens, D. 2015. Asymmetry in genitalia does not increase the rate of their evolution. Molecular phylogenetics and evolution 93: 180–187.
• Özgül-Siemund, A. & Ahrens, D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to zoology 84 (2): 167–178.

2016

• Ahrens, D. & Fabrizi, S. 2016. A monograph of Sericini of India (Coleoptera: Scarabaeidae). Bonn zoological bulletin 65(1 & 2): 1–355. Full article Reference page. 
• Ahrens, D., Fabrizi, S. & Rößner, E. 2016. A new species of Maladera from Jordan (Coleoptera: Scarabaeidae: Sericini). Zoology in the Middle East 62(4): 347–351. DOI: 10.1080/09397140.2016.1250704 . Reference page. 
• Liu, W-G., Fabrizi, S., Bai, M. & Ahrens, D. 2016. A taxonomic revision of Neoserica (sensu lato): the species groups N. lubrica, N. obscura, and N. silvestris (Coleoptera, Scarabaeidae, Sericini). Zookeys, 635: 123–160. DOI: 10.3897/zookeys.635.9915. Reference page. 

2017

• Li, W-J., Huo, L-Z., Ahrens, D., Ren, S-X. & Wang, X-M. 2017. Renius cornutus, a new genus and species of Chilocorini from Tibet, China (Coleoptera, Coccinellidae). ZooKeys 678: 121—128. DOI: 10.3897/zookeys.678.11862. Reference page. 
• Liu, W-G., Bai, M., Yang, X.K. & Ahrens, D. 2017d. New species and records of Lasioserica and Gynaecoserica from China (Coleoptera, Scarabaeidae, Sericini). Bonn zoological Bulletin, 66(1): 29–36. full article (PDF). Reference page. 
• Liu, W-G., Fabrizi, S., Yang, X-K., Bai, M. & Ahrens, D. 2017. New species of Nipponoserica and Paraserica from China (Coleoptera, Scarabaeidae, Sericini). ZooKeys 721: 65–91. DOI: 10.3897/zookeys.721.13918. Reference page. 

2018

• Sreedevi, K., Speer, J., Fabrizi, S. & Ahrens, D. 2018. New species and records of Sericini scarab beetles from the Indian subcontinent (Coleoptera, Scarabaeidae). ZooKeys 772: 97–128. DOI: 10.3897/zookeys.772.25320 Reference page. 

2019

• Ahrens, D., Fabrizi, S., Nikolaev, K., Knechtges, L. & Eberle, J. 2019. On the identity of some taxa of Sericinae described by C. P. Thunberg and L. Gyllenhal (Coleoptera, Scarabaeidae). Bonn zoological bulletin, 68(1): 21–29. DOI: 10.20363/BZB-2019.68.1.021. . Reference page. 
• Fabrizi, S., Dalstein, V. & Ahrens, D. 2019. A monograph on the genus Tetraserica from the Indochinese region (Coleoptera, Scarabaeidae, Sericini). Zookeys, 837: 1–155. DOI: 10.3897/zookeys.837.32057 Reference page. 
• Fabrizi, S., Eberle, J. & Ahrens, D. 2019. Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society, 186(4): 950–982. DOI: 10.1093/zoolinnean/zly095 . Reference page. 
• Sreedevi, K., Ranasinghe, U.G.S.L., Fabrizi, S. & Ahrens, D. 2019. New species and records of Sericini from the Indian subcontinent (Coleoptera, Scarabaeidae) II. European Journal of Taxonomy 567: 1–26. DOI: 10.5852/ejt.2019.567. . Reference page. 

2020

• Ahrens, D. & Pacheco, T.L. 2020. Maladera rugosa (Blanchard, 1850) new combination—a valid species and senior synonym of Maladera graeca Petrovitz, 1969 (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 4759(3): 443–445. DOI: 10.11646/zootaxa.4759.3.12. . Reference page. .
• Pacheco, T.L., Monné, M.L., Vaz-de-Mello, F.Z. & Ahrens, D. 2020. Notes on the taxonomy of some "Astaena" species described by Burmeister (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 4885(1): 143–145. DOI: 10.11646/zootaxa.4885.1.11 Reference page. 

2021

• Ahrens, D., Liu, W.G., Pham, V.P. & Fabrizi, S. 2021. An overview on the genus Amiserica Nomura, 1974 (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5050(1): 001–063. DOI: 10.11646/zootaxa.5050.1.1. . Reference page. 
• Bhunia, D., Gupta, D., Chandra, K. & Ahrens, D. 2021. New species and records of Sericini of India (Coleoptera: Scarabaeidae: Melolonthinae) II. Zootaxa 5081(4): 594–600. DOI: 10.11646/zootaxa.5081.4.10. . Reference page. 
• Fabrizi, S., Liu, W.G., Bai, M., Yang, X.K. & Ahrens, D. 2021. A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 4922(1): 001–400. DOI: 10.11646/zootaxa.4922.1.1. . Reference page. 
• Pacheco, T.L., Monné, M.L., Vaz-de-Mello, F.Z. & Ahrens, D. 2021. Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L.W. Saylor. European Journal of Taxonomy 750(1): 94–123. DOI: 10.5852/ejt.2021.750.1365 . Reference page. 

2022

• Ahrens, D. & Lukic, D. 2022. Neoserica (s. str.) phuphami—a further new Neoserica species from Vietnam with highly modified pronotum (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5104(3): 441–444. DOI: 10.11646/zootaxa.5104.3.8. . Reference page. 
• Pacheco, T.L., Monné, M.L., Vaz-de-Mello, F.Z. & Ahrens, D. 2022. First non‑feeding Sericini beetles (Coleoptera, Scarabaeidae): new genus from Amazonia and phylogenetic position. Organisms Diversity & Evolution 22(3): 733–748. DOI: 10.1007/s13127-022-00555-x . Reference page. 
• Pacheco, T.L., Wipfler, B., Monné, M.L. & Ahrens, D. 2022. The genus Symmela Erichson, 1835 (Coleoptera, Scarabaeidae, Sericini): taxonomy and phylogenetic analysis. Journal of Natural History 56 (9–12): 607–705. DOI: 10.1080/00222933.2022.2084649 . Reference page. 
• Pacheco, T.L., Wipfler, B., Monné, M.L. & Ahrens, D. 2022. The genus Raysymmela Saylor, 1947 (Coleoptera, Scarabaeidae, Melolonthinae, Sericini): taxonomy and phylogenetic analysis. Insect Systematics and Evolution 53(4): 400–441. DOI: 10.1163/1876312X-bja10031 . Reference page. 
• Pacheco, T.L., Monné, M.L., Vaz-de-Mello, F.Z. & Ahrens, D. 2022. First non‑feeding Sericini beetles (Coleoptera, Scarabaeidae): new genus from Amazonia and phylogenetic position. Organisms Diversity & Evolution 22(3): 733–748. DOI: 10.1007/s13127-022-00555-x . Reference page. 
• Pacheco, T.L., Monné, M.L. & Ahrens, D. 2022. Comparative analysis of morphospace of Neotropical Sericini (Coleoptera: Scarabaeidae): disparity in the light of species diversity and activity patterns. Organisms Diversity & Evolution 22(1): 177–188. DOI: 10.1007/s13127-021-00530-y . Reference page. 

2023

• Ahrens, D., Liu, W.G., Lukic, D. & Bai, M. 2023. A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241(1): 001–115. DOI: 10.11646/zootaxa.5241.1.1. . Reference page. 
• Ahrens, D., Lukic, Pham, P., Li, W. & Liu, W-G. 2023. Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species, and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini). Zootaxa 5374(4): 451–486. DOI: 10.11646/zootaxa.5374.4.1. . Reference page. 
• Pacheco, T.L. & Ahrens, D. 2023. Addendum to the revision of the South American genus Symmela Erichson, 1835 (Coleoptera: Scarabaeidae: Melolonthinae: Sericini): a further new species from Brazil. Zootaxa 5263(2): 297–300. DOI: 10.11646/zootaxa.5263.2.8 Reference page. 
• Pacheco, T.L., Monné, M.L. & Ahrens, D. 2023. Morphology-based phylogenetic analysis of South American Sericini chafers (Coleoptera, Scarabaeidae) contrasts patterns of morphological disparity and current classification. Zoologischer Anzeiger 302: 43–57. DOI: 10.1016/j.jcz.2022.11.004 . Reference page. 

External links

• Zoologisches Forschungsmuseum Alexander Koenig