Brett Charles Ratcliffe

Brett Charles Ratcliffe Entomologist, USA.

  • Curator & Professor Systematics Research Collections and Department of Entomology, University of Nebraska, U.S.A.
  • Systematics Research Collections, University of Nebraska State Museum, University of Nebraska, Lincoln, Nebraska, 68588-0338, United States of America.
  • E-mail:
  • Specialist in Scarabaeidae.

Taxon names authored

(List may be incomplete)



(List may be incomplete)










  • Ratcliffe, B.C. 1976: A revision of the genus Strategus (Coleoptera: Scarabaeidae). Bulletin of the University of Nebraska State Museum, 10: 93–204, 176 figs.
  • Ratcliffe, B.C. 1976: Notes on the biology of Euphoriaspis hirtipes (Horn) with a description of the larva and pupa (Coleoptera: Scarabaeidae). Coleopterists bulletin, 30: 217–225.
  • Pike, K.S., R.L. Rivers, Ratcliffe, B.C.; C.Y. Oseto; Z B Mayo, 1976: A world bibliography of the genus Cyclocephala (Coleoptera: Scarabaeidae). University of Nebraska Agricultural Experiment Station Miscellaneous Publication, 32: 1–32.


  • Ratcliffe, B.C. 1977. A case of tarantula induced vesicular dermatitis. Journal of Medical Entomology, 13: 745–747.
  • Ratcliffe, B.C. 1977. A description of the larva and pupa of Osmoderma subplanata Casey and Cremastocheilus wheeleri LeConte (Coleoptera: Scarabaeidae). Journal of the Kansas Entomological Society, 50: 363–370.
  • Ratcliffe, B.C. 1977. Four new species of Neotropical Cyclocephalini (Coleoptera: Scarabaeidae). Acta amazonica, 7: 429–434.


  • Ratcliffe, B.C. 1978. The systematic entomology collections of Brazil's National Institute for Amazonian Research (INPA). Bulletin of the Entomological Society of America, 24: 62.
  • Ratcliffe, B.C. 1978. As coleções de entomologia sistematica do Instituto Naçional de Pesquisas da Amazonia (INPA-Manaus). Ciencia e Cultura, 30: 627–628.
  • Ratcliffe, B.C. 1978. A review of the genus Argyripa (Coleoptera: Scarabaeidae). Systematic Entomology 3: 371–378.
  • Ratcliffe, B.C. 1978. A new species of Trox from the Amazon Basin with new distribution records for Central American Trox (Coleoptera: Scarabaeidae). Acta amazonica, 8: 299–302.
  • Ratcliffe, B.C. 1978. New species of Stenocrates from Brazil (Coleoptera: Scarabaeidae). Acta amazonica, 8: 489–495.
  • Ratcliffe, B.C.; N. D. Penny, 1978. Catalogo dos tipos de invertebrados depositados nas coleções entomologicas do INPA-Manaus. Acta amazonica, 8: 689–697.


  • Ratcliffe, B.C.; J.A. Fagerstrom, 1980. Invertebrate Lebensspuren of Holocene floodplains: their morphology, origin and paleoecological significance. Journal of Paleontology, 54: 614–630.
  • Ratcliffe, B.C.; F. Chalumeau, 1980. Strategus syphax (Fabr.): A description of the third instar larva and pupa (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 34: 85–93.


  • Ratcliffe, B.C. 1981. New species of Coprini (Coleoptera: Scarabaeidae: Scarabaeinae) taken from the pelage of three toed sloths (Bradypus tridactylus L.) in central Amazonia with a commentary on scarab-sloth relationships. Coleopterists bulletin, 34: 337–350.
  • Ratcliffe, B.C. 1981. The fourth Hylobothynus: spanning the Amazonian gap (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 35: 303–306.
  • Ratcliffe, B.C. 1981. New species and distribution records of Surutu from Amazonian Brazil (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 35: 107–112.
  • Ratcliffe, B.C. 1981. A redescription of Trichillum hirsutum Boucomont with notes on other interesting Brazilian Scarabaeinae (Coleoptera: Scarabaeidae). Acta Amazonica, 11: 183–186.
  • Ratcliffe, B.C. 1981. Barutus hartmanni, a new genus and species from Panama with a key to the New World genera of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 35: 463–472.
  • Ratcliffe, B.C. 1981. Xyloryctes jamaicensis (Drury) in Nebraska. Scarabaeus, 4: 8.


  • Ratcliffe, B.C. 1982. American Oryctini: Strategus verrilli Ratcliffe rediscovered and described, and new records and comments for other Strategus and Hispanioryctes (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 36: 352–357.
  • Ratcliffe, B.C. 1982. Barutus hartmanni, a new genus and species from Panama with a key to the genera of New World Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin 35(4): 463–472. JSTOR


  • Ratcliffe, B.C. 1983. Reprinted in Sarjeant, William A.S. (Ed.), Terrestrial Trace Fossils. Benchmark Papers in Geology 76, pp. 251–267. Hutchinson Ross Publishing Company, Stroudsburg, Pa.
  • Ratcliffe, B.C. 1983. Trox hamatus Robinson (Troginae) using a Canthon (Scarabaeinae) brood ball and new records of North American Trox (Coleoptera: Scarabaeidae). Transactions of the Nebraska Academy of Sciences, 11: 53–55.
  • Ratcliffe, B.C.; R.-P. Dechambre, 1983. New combinations, synonymy and distribution records for Neotropical Pentodontini and Oryctini (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 37: 267–272.
  • Morón, M. ; Ratcliffe, B.C., 1984. Description of the larva and pupa of Argyripa lansbergei (Sallé) with new distributional records for the genus and a key to New World Gymnetini larvae (Coleoptera: Scarabaeidae: Cetoniinae). Proceedings of the Entomological Society of Washington, 86: 760–768.


  • Ratcliffe, B.C. 1984. A review of the Penichrolucaninae with analyses of phylogeny and biogeography and description of a second New World species from the Amazon Basin (Coleoptera: Lucanidae). Quaestiones Entomologique 20: 60–87.


  • Ratcliffe, B.C. 1985. The Museum für Naturkunde, Berlin. Scarabaeus No. 11: 1–3.
  • Penny, N. D.; Ratcliffe, B.C., 1985. Entomological collections and human resources in Brazil. ASC Newsletter 13: 21–24.
  • Boos, J.; Ratcliffe, B.C., 1985. A new subspecies of Inca clathrata (Olivier) from Trinidad, West Indies, and range extensions for Inca clathrata sommeri Westwood (Coleoptera: Scarabaeidae: Trichiinae). Coleopterists bulletin, 39: 381–389.
  • Ratcliffe, B.C. 1985. God is, forsooth, a scarab: a rebuttal to Acorn’s repugnant carabid hypothesis. Biologie und Naturwissenschaften der Käfer 3: 9–19.






  • Ratcliffe, B.C.; M. L. Jameson, 1989. A synopsis of the genus Areoda (Coleoptera: Scarabaeidae: Rutelinae). Coleopterists bulletin, 43: 135–144.
  • Morón, M.; Ratcliffe, B.C., 1989. A synopsis of the American Goliathini with description of a new Neoscelis from Mexico (Coleoptera: Scarabaeidae: Cetoniinae). Coleopterists bulletin, 43: 339–348.
  • Ratcliffe, B.C.; G. Clark, 1989. Observations on the tunnel morphology of Heterocerus brunneus Melsh. (Coleoptera: Heteroceridae) and the paleoecological significance of insect traces. Journal of Paleontology, 63: 228–232.
  • Ratcliffe, B.C. (Editor), 1989. University of Nebraska State Museum. ASC Newsletter, 17: 1, 3, 10–12.
  • Ratcliffe, B.C.; M. L. Jameson, 1989. A new species of Pelidnota (Coleoptera: Scarabaeidae: Rutelinae) from Panama. Coleopterists bulletin, 43: 259–262.
  • Ratcliffe, B.C. 1989. A case of gynandromorphy in Golofa tersander (Coleoptera: Scarabaeidae). Coleopterists bulletin, 43: 256–258.
  • Ratcliffe, B.C. 1989. Corrections and clarifications to Endrödi’s “The Dynastinae of the World.” Coleopterists bulletin, 43: 275–278.


  • Ratcliffe, B.C.; L. Delgado, 1990. New species and notes of Cyclocephala from Mexico (Coleoptera: Scarabaeidae: Dynastinae). Folia Entomologica Mexicana No. 80: 41–57.
  • Ratcliffe, B.C. 1990. The significance of scarab beetles in the ethnoentomology of non-industrial, indigenous peoples. Proc. I Internat. Congr. Ethnobiology, Belem, Brazil, vol. 1: 159–185.
  • Morón, M.; Ratcliffe, B.C., 1990. Descriptions of Strategus larvae with a new key to species based on the larvae (Coleoptera: Scarabaeidae: Dynastinae). Elytron 4: 53–66.
  • Ratcliffe, B.C. 1990. Prairie scarabs. The Nebraska perspective. American Entomologist 36: 28–35.
  • Howden, H. F.; Ratcliffe, B.C., 1990. An unusual new species of Trigonopeltastes Burmeister from Chiapas, Mexico (Coleoptera: Scarabaeidae: Trichiinae). Canadian Entomologist 122: 1–4.
  • Ratcliffe, B.C. 1990. An unusual method of pattern disruption in an Amazonian scarab beetle. Boletim Museu Paraense Emilio Goeldi, Series Zoologia 6:109–113.


  • Ratcliffe, B.C. 1991. The Passalidae and Lucanidae (Insecta: Coleoptera) of Nebraska. Great Plains Research 1: 249–282.


  • Ratcliffe, B.C. 1992. New species and country records of Brazilian Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae). Tijdschrift voor Entomologie 136: 179–190. Reference page
  • Ratcliffe, B.C. 1992a. Nine new species and 11 country records of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Panama and Costa Rica. The Coleopterists Bulletin 46: 216–235. Reference page
  • Ratcliffe, B.C. 1992. Two new species of Cyclocephala from Arizona and Mexico and a note on melanistic C. melanocephala (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 46: 250–255.
  • Ratcliffe, B.C.; C. Deloya, 1992. The biogeography and phylogeny of Hologymnetis (Coleoptera: Scarabaeidae: Cetoniinae) with a revision of the genus.Coleopterists bulletin, 45: 161–202.
  • {{aut|Ratcliffe, B.C. 1992. A new species of Ancognatha from Panama (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 46: 256–259.
  • Ratcliffe, B.C.; M. L. Jameson, 1992. New Nebraska occurrences of the endangered American burying beetle (Coleoptera: Silphidae). Coleopterists bulletin, 46: 421–425.
  • Ratcliffe, B.C., M. L. Jameson, and T. Taylor, 1992. Two remarkable new species of Plusiotis (Coleoptera: Scarabaeidae: Rutelinae) from Mexico and Central America. Insecta mundi, 6: 1–5.
  • Ratcliffe, B.C.; C. L. Messenger, 1992. Vial support system for fluid collections, pp. 213–214. In: C. Rose and A.R. de Torres (eds.), Storage of Natural History Collections: Ideas and Practical Solutions. Society for the Preservation of Natural History Collections, Pittsburgh, PA.


  • Jameson, M. L., B. C. Ratcliffe, and M. M. Morón, 1994. A synopsis of the Neotropical genus Calomacraspis Bates with a key to larvae of the American genera of Rutelini (Coleoptera: Scarabaeidae: Rutelinae). Annals of the Entomological Society of America 87: 43–58.
  • Ratcliffe, B.C. 1994. Orizabus subaziro, a new species from Mexico (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 48: 91–93.


  • Ratcliffe, B.C. 1995. American burying beetle. Nebraska’s Threatened and Endangered Species insert in NEBRASKA land Magazine 73, 6 pp.


  • Maes, J.-M.; Ratcliffe, B.C., 1996. Scarabaeidae nuevos para la fauna de Nicaragua. Revista Nicaraguense de Entomologia 34: 17–18.


  • Ratcliffe, B.C.; M. A. Morón, 1997. Capítulo 3. Dynastinae, pp. 53–98. In Morón, Miguel A., Brett C. Ratcliffe, and Cuauhtemoc Deloya, Atlas de los Escarabajos de Mexico (Coleoptera Lamellicornia). Vol. 1. Familia Melolonthidae. CONABIO and Sociedad Mexicana de Entomologia, Mexico. 280 pp.
  • Maes, J.-M., B. C. Ratcliffe, and M. L. Jameson, 1997. Fauna entomologica de la Reserva Natural BOSAWAS, Nicaragua. XI. Escarabajos (Coleoptera: Scarabaeidae) nuevos para la fauna de Nicaragua. Revista Nicaraguense de Entomologia 39: 41–45.
  • Morón, M. A.; Ratcliffe, B.C., 1997. New tribal placement of the genus Coscinocephalus Prell 1936, with description of the larva, pupa, and adult of a new species from Mexico (Coleoptera: Scarabaeoidea: Dynastinae). Journal of the New York Entomological Society 104: 48–61.


  • Ratcliffe, B.C. 1998. An unusual new unicorn species of Copris from Nicaragua (Scarabaeidae: Scarabaeinae, Coprini. Coleopterists bulletin, 52: 93–96.
  • Ratcliffe, B.C. 1998. Insects., pp. 143–154. In Bleed, A. S. and C. A. Flowerday (eds.), An Atlas of the Sand Hills. Resource Atlas 5b, Conservation and Survey Division, Institute of Agriculture and Natural Resources, University of Nebraska-Lincoln. 260 pp.
  • Ratcliffe, B.C.; M. A. Ivie, 1998. New species of Hemiphileurus (Coleoptera: Scarabaeidae: Dynastinae) from the Dominican Republic with a key to the West Indian species of Hemiphileurus. Coleopterists bulletin, 52: 201–208.


  • Ratcliffe, B.C.; A. B. T. Smith, 1999. New species of Canthonella Chapin (Scarabaeidae: Scarabaeinae) from Amazonian Brazil. Coleopterists bulletin, 53: 1–7.
  • Bedick, J. C., B. C. Ratcliffe, W. W. Hoback, and L. G. Higley, 1999. Distribution, ecology, and population dynamics of the American burying beetle [Nicrophorus americanus Olivier (Coleoptera, Silphidae)] in south-central Nebraska, USA. Journal of Insect Conservation 3: 171–181.
  • Ratcliffe, B.C. 1999. U. S. scarab collection moves to Nebraska. ASC Newsletter 27: 1 and 6.
  • Coca-Abia, M., P. Dominguez-Alonso,; Ratcliffe, B.C., 1999. Evidencias de actividad biológica producidas por artropodos terrestres a lo largo del tiempo geológico, pp. 213–221. In Melic, A., J. J. de Haro, M. Mendez, and I. Ribera (Eds.), Evolución y Filogenia de Arthropoda. Boletin de la Sociedad Entomológica Aragonesa (Monographic Volume) 26: 1–806.


  • Ratcliffe, B.C. 2000. Dynastine scarab beetles of Monteverde, pp. 111–113. In Nadkarni, N. and N. Wheelwright (eds.), Monteverde. Ecology and Conservation of a Tropical Cloud Forest. Oxford University Press, New York, NY. 573 pp.


  • Mico, E., W. E. Hall,; Ratcliffe, B.C., 2001. Descriptions of the larvae of Hoplopyga singularis (Gory and Percheron) and Hologymnetis cinerea (Gory and Percheron) with a revised key to the larvae of New World Gymnetini (Coleoptera: Scarabaeidae: Cetoniinae). Coleopterists bulletin, 55: 205–217.
  • Ratcliffe, B.C.; E. Mico, 2001. A review of the Neotropical genus Neocorvicoana Ratcliffe and Micó, new genus (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini). Coleopterists bulletin, 55: 279–296.
  • Ratcliffe, B.C.; F. Ocampo, 2001. Tyrannasorus rex Ratcliffe and Ocampo, a new genus and species of Miocene hybosorid in amber from the Dominican Republic (Coleoptera: Scarabaeoidea: Hybosoridae). Coleopterists bulletin, 55: 351–355.
  • Ratcliffe, B.C. 2001. New species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) from Mexico, Guatemala, Colombia, and Brazil. Coleopterists bulletin, 55: 433–443.


  • Ratcliffe, B.C. 2002. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32: 1–48. Abstract & excerpt Reference page
  • Ratcliffe, B.C. & J. Christen 2002. Lucanus elaphus Fabr. (Coleoptera: Lucanidae) discovered in Nebraska, with brief notes on its biology and identification. Entomological News 113: 68–70.
  • Ratcliffe, B.C. & R. D. Cave, 2002. New species of Cyclocephala from Honduras and El Salvador (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini). Coleopterists bulletin, 56: 152–157.
  • Ratcliffe, B.C. & Ocampo, F.C. 2002: A review of the genus Hylamorpha Arrow (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini: Brachysternina). The Coleopterists Bulletin 56(3): 367–378. DOI: 10.1649/0010-065X(2002)056[0367:AROTGH]2.0.CO;2 Reference page
  • Ratcliffe, B.C. 2002. Review of the genus Palaeophileurus (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with description of two new species from Peru. Annals of the Entomological Society of America 95: 335–339.
  • Ratcliffe, B.C. & P. C. Hammond 2002. Insects and the native vegetation of Nebraska. Transactions of the Nebraska Academy of Sciences 28: 29–47.
  • Jameson, M. L., Ratcliffe, B.C. & V. Maly, 2002. Review of the genus Acrobolbia Ohaus with remarks on its classification and key to the world genera of Cyclocephalini (Coleoptera: Scarabaeidae: Dynastinae). Folia Heyrovskyana 10: 1–15.
  • Jameson, M. L. & Ratcliffe, B.C. 2002. Series Scarabaeiformia Crowson 1960, Superfamily Scarabaeoidea Latreille 1802, pp. 1–5. In, Arnett, R. H., M. Thomas, P. E. Skelley, and J. H. Frank (eds.), American Beetles, Volume 2. CRC Press, Boca Raton, FL. 861 pp.
  • Ratcliffe, B.C. 2002. Chapter 23. Lucanidae Latreille 1804, pp. 6–9. In, Arnett, R. H., M. C. Thomas, P. E. Skelley, and J. H. Frank (eds.), American Beetles, Volume 2. CRC Press, Boca Raton, FL. 861 pp.
  • Jameson, M. L. & Ratcliffe, B.C. 2002. Chapter 24. Diphyllosomatidae Holloway 1972, pp. 10–11. In, Arnett, R. H., M. C. Thomas, P. E. Skelley, and J. H. Frank (eds.), American Beetles, Volume 2. CRC Press, Boca Raton, FL. 861 pp.
  • Ratcliffe, B.C., M. L. Jameson, and A. B. T. Smith, 2002. Chapter 34. Scarabaeidae Latreille 1802, pp. 39–81 (in part). In, Arnett, R. H., M. C. Thomas, P. E. Skelley, and J. H. Frank (eds.), American Beetles, Volume 2. CRC Press, Boca Raton, FL. 861 pp.
  • Ratcliffe, B.C. 2002. Chapter 34-V. Dynastinae MacLeay 1819, pp. 64–67. In, Arnett, R. H., M. C. Thomas, P. E. Skelley, and J. H. Frank (eds.), American Beetles, Volume 2. CRC Press, Boca Raton, FL. 861 pp.
  • Ratcliffe, B.C. 2002. Chapter 34-VI. Cetoniinae Leach 1815, pp. 67–71. In, Arnett, R. H., M. C. Thomas, P. E. Skelley, and J. H. Frank (eds.), American Beetles, Volume 2. CRC Press, Boca Raton, FL. 861 pp.


  • Ratcliffe, B.C. 2003. New species of Hemiphileurus Kolbe from Honduras and Guatemala (Coleoptera: Scarabaeidae: Dynastinae: Phileurini). Coleopterists bulletin, 57: 334–338.
  • Ratcliffe, B.C. 2003. The Dynastine Scarab beetles of Costa Rica and Panama. Bulletin of the University of Nebraska State Museum 16: 1–506.
  • Mudge, A. D., B. C. Ratcliffe, R. L. Westcott, and F. A. Noguera F.A., 2003. A new species of Neoscelis from Jalisco, Mexico (Coleoptera: Scarabaeidae: Cetoniinae). Folia Heyrovskyana 11: 143–154.


  • Ratcliffe, B.C. 2004. Comment on the proposed precedence of Bolboceras Kirby, 1819 (July) (Insecta, Coleoptera) over Odonteus Samouelle, 1819 (June). Bulletin of zoological nomenclature, 61: 43.
  • Bedick, J., B. C. Ratcliffe, and L. G. Higley, 2004. A new sampling protocol for the American burying beetle, Nicrophorus americanus Olivier (Coleoptera: Silphidae). Coleopterists bulletin, 58: 57–70.
  • Ratcliffe, B.C.; M. L. Jameson, 2004. The revised classification for Scarabaeoidea: what the hell is going on? Scarabs, 15: 3–10.
  • Ratcliffe, B.C. 2004. Lectotype designations in the New World Gymnetini (Coleoptera: Scarabaeidae: Cetoniinae). Zootaxa, 729: 1–19.




  • Ratcliffe, B.C. 2006. VII Reunión Latinoamericana de Scarabaeoidología. Scarabs, 19: 1–9.
  • Ratcliffe, B.C. 2006. Heterogomphus effeminatus, an unusual new species of rhinocerosbeetle from French Guiana (Coleoptera: Scarabaeidae: Dynastinae: Oryctini). Acta Zoologica Cracoviensia 49b: 9–12.
  • Ratcliffe, B.C. 2006. Scarab beetles in human culture, pp. 85–101. In, Jameson, M. L.; Ratcliffe, B.C. (editors), Scarabaeoidea in the 21st Century: a Festschrift Honoring Henry F. Howden. The Coleopterists Society Monograph 5: 1–216.
  • Neita, J. C., J. Orozco,; Ratcliffe, B.C., 2006. Escarabajos (Scarabaeidae: Pleurosticti) de la selva baja del bosque pluvial tropical “BP-T”, Chocó, Colombia. Acta zoológica mexicana (n. s.), 22: 1–32.


  • vUtz, R. M., B. C. Ratcliffe, B. T. Moore, and K. J. Hartman, 2007. Disproportionate relative importance of a terrestrial beetle family (Coleoptera: Scarabaeidae) as a prey source for central Appalachian brook trout. Transactions of the American Fisheries Society, 136: 177–184.
  • Ratcliffe, B.C. 2007. A remarkable new species of Megaceras from Peru (Coleoptera: Scarabaeidae: Dynastinae). The “Dim Effect”: nature mimicking art, Coleopterists bulletin, 61: 463–467.
  • Neita Moreno, J. C., B. C. Ratcliffe, and G. Calberto, 2007. Immature stages of Aspidolea singularis (Scarabaeidae: Dynastinae: Cyclocephalini). Revista Colombiana de Entomología 33: 178–182.
  • Ratcliffe, B.C.; A. B. T. Smith, 2007. Bothynus Hope, 1837 (Insecta, Coleoptera, Scarabaeidae): proposed conservation of usage by designation of Scarabaeus ascanius Kirby, 1819 as the type species. Bulletin of zoological nomenclature, 64:29–31


  • Ratcliffe, B.C.; R. D. Cave, 2008. A biotic survey and inventory of the dynastine scarab beetles of Central America, North America, and the Caribbean regions: review of a long term, multi-country project. Zoosystema 30: 651–663.
  • Ratcliffe, B.C.; R. D. Cave, 2008. The Dynastinae (Coleoptera: Scarabaeidae) of the Bahamas with a description of a new species of Cyclocephala from Great Inagua Island. Insecta mundi, 0024: 1–10.
  • Gasca, H. J., C. R. Vasconcelos da Fonseca,; Ratcliffe, B.C., 2008. Synopsis of the Oryctini (Coleoptera: Scarabaeidae: Dynastinae) from the Brazilian Amazon. Insecta mundi, 0061: 1–62.
  • Genoways, H. H.; Ratcliffe, B.C., 2008. Engineer Cantonment, Missouri Territory, 1819–1820: America’s First Biodiversity Inventory. Great Plains Research 18: 3–31.
  • Ratcliffe, B.C. 2008. More new species of Cyclocephala from South America (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 62: 221–241.
  • Cave, R. D.; Ratcliffe, B.C., 2008. Scarab beetles (Coleoptera: Scarabaeoidea), pp. 3273–3286. In, J. Capinera (Editor), Encyclopedia of Entomology, 2nd Edition. Springer Netherlands. 4000 pp.
  • Santiago-Blay, J. A., B. C. Ratcliffe, F.-T. Krell, and R. Anderson, 2008. Allotypes should be from the type series: a position paper for reinstating Recommendation 72A from the third edition of the Code that defines the term “allotype”. Bulletin of zoological nomenclature, 65: 260–264.


  • Jacobs, J., I. Noble, S. Palminteri,; Ratcliffe, B.C., 2009 (2008). First come, first serve: “sit and wait” behavior in dung beetles at the source of primate dung. Neotropical entomology, 37: 641–645.
  • Ratcliffe, B.C. 2009. A most unusual scarab. Unusual (en / yoo / zh / ooel): unlike what is expected, differing in some way from the norm. Weird. Scarabs 40: 11–14.
  • Jameson, M. L.; Ratcliffe, B.C., 2009. Revision of the genus Chalcasthenes Arrow (Scarabaeidae: Dynastinae: Oryctoderini) from the Solomon Islands. Journal of the Australian Entomological Society 48: 149–163.
  • Cave, R. D.; Ratcliffe, B.C., 2009. Sondeo biótico e inventario de los escarabajos dynástinos de Mesoamerica, Norteamerica y las Antillas: un proyecto multinacional a largo plazo. Cuadernos de Biodiversidad No. 29: 5–13.
  • Ratcliffe, B.C. 2009. Xyloryctes (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) in the United States. Qui es et ubi fuisti et quo vadis? Insecta mundi, 0100:1–11.
  • Ratcliffe, B.C.; R. D. Cave, 2009. New species of Cyclocephala Dejean, 1821 from Guatemala (Scarabaeidae: Dynastinae: Cyclocephalini). Coleopterists bulletin, 63: 325–332.
  • Neita, J. C.; Ratcliffe, B.C., 2009. Description of the adult female, larva, pupa, and distribution of Strategus fascinus Burmeister (Scarabaeidae: Dynastinae: Oryctini). Coleopterists bulletin, 63: 367–379.
  • Ratcliffe, B.C. & Orozco, J. 2009: A review of the biology of Pericoptus truncatus (Fabr.) (Coleoptera: Scarabaeidae: Pentodontini) from New Zealand and a revised description of the third instar. The Coleopterists Bulletin 63(4): 445–451. DOI: 10.1649/1186.1 Reference page
  • Jameson, M. L., D. E. Oishi, B. C. Ratcliffe, and G. T. McQuate., 2009. Two additional invasive scarabaeoid beetles (Coleoptera: Scarabaeidae: Dynastinae) in Hawaii. Proceedings of the Hawaiian Entomological Society 41: 25–30.


  • Ratcliffe, B.C. 2010. A review of the Blaesiina (Coleoptera, Scarabaeidae, Cetoniinae, Gymnetini). Zookeys 34: 105–128.
  • Ratcliffe, B.C. & J.-P. Saltin, 2010. Melanistic Mitracephala humboldti Thomson (Coleoptera: Scarabaeidae: Dynastinae). A tale from the dark side. Scarabs 51: 1–9.
  • Céspedes, A.; Ratcliffe, B.C., 2010. Golofa clavigera (Linnaeus, 1771) in Bolivia: a new country record (Coleoptera: Scarabaeidae: Dynastinae). Ecología en Bolivia (Revista del Instituto de Ecología, Carrera de Biología, Facultad de Ciencias Puras y Naturales, Universidad Mayor de San Andrés, La Paz, Bolivia) 45: 73–76.
  • Ratcliffe, B.C. 2010. A new Bolivian Bothynus Hope, 1837, with a key to the Bolivian species (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Coleopterists bulletin, 64: 105–108.
  • Ratcliffe, B.C. & Cave, R.D. 2010. New species of Orizabus (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) from Mexico and Guatemala with a revised key and checklist of the species in the genus. Insecta mundi, 0127: 1–16.
  • Ratcliffe, B.C.; A. Zubair, 2010. Additions to the distribution of Scarabaeidae (Insecta: Coleoptera) in northern Pakistan. Pakistan Journal of Zoology 42: 827–830.
  • Ratcliffe, B.C. & Cave, R.D. 2010. The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with a description of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini) from Little Cayman. Insecta mundi, 0139: 1–15. PDF
  • Ratcliffe, B.C.; J.-P. Saltin, 2010. Homophileurus neptunus Dechambre, a new junior synonym of H. waldenfelsi Endrödi (Coleoptera: Scarabaeidae: Dynastinae: Phileurini). Insecta mundi, 0146: 1–5.
  • Ratcliffe, B.C. 2010. Gorditus rotundus, a new genus and species of Pentodontini from Cuba (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa 2683: 56–60. Preview
  • Neita, J. C.; Ratcliffe, B.C., 2010. A new species of Hemiphileurus Kolbe from Colombia, with description of its larva and pupa (Coleoptera: Scarabaeidae: Dynastinae: Phileurini). Coleopterists bulletin, 64: 347–355.


  • Ratcliffe, B.C. & Cave, R.D. 2011. A revision of the genera Endroedianibe Chalumeau and Hispanioryctes Howden and Endrödi from Hispaniola with descriptions of new species (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin 65: 1–14.
  • Ratcliffe, B.C. 2011. Hoplopygothrix Schürhoff revisited: a new species and country record for Bolivia (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini). Coleopterists bulletin 65: 63–66. DOI: 10.1649/0010-065X-65.1.63
  • Ratcliffe, B.C. 2011. A review of the species of Phileurus with quadridentate protibiae and description of a new species from Panama (Coleoptera: Scarabaeidae: Dynastinae: Phileurini). Coleopterists bulletin, 65: 125–129.
  • Ratcliffe, B.C. & Warner, W. B. 2011. A revision of the genus Gymnetina Casey, 1915 (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini). Insecta mundi, 0173: 1–28.
  • Ratcliffe, B.C. & Curoe, D., 2011. 2011: A New Species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) from Panama, with a Revised Key to the Panamanian Species. Coleopterists bulletin, 65(2): 139–143. Abstract: DOI: 10.1649/072.065.0208.
  • Neita, J.C. & Ratcliffe, B.C. 2011. Immature stages of Homophileurus tricuspis Prell (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with notes on natural history and distribution in Colombia. Zootaxa 2915: 20–28.
  • Ratcliffe, B.C. & Hoffman, R.L., 2011. Cyclocephala nigricollis Burmeister. What is it? And where is it? (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini). Coleopterists bulletin 65: 135–138.
  • Gasca, H.J. & Ratcliffe, B.C. 2011. An annotated checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms. Zootaxa 3090: 21–40.
  • Ratcliffe, B.C. & Skelley, P. 2011. Description of the larva of Homophileurus integer (Burmeister, 1847) (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with a key to Homophileurus larvae and notes on biology. Coleopterists bulletin, 65: 297–304.


  • Saltin, J.-P. & Ratcliffe, B.C. 2012. A review of the distribution of Harposcelis paradoxus Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) with a new country record for Peru. Dugesiana, 18: 147–151 and 19: 10 (correction).
  • Ratcliffe, B.C. 2012. First Reported Occurrence of Hoplopyga Thomson, 1880 (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini) from the West Indies, with Description of a New Species. Coleopterists bulletin 66(2): 111–115. DOI: 10.1649/072.066.0203
  • Jacobs, J. M., R. von May & Ratcliffe, B.C. 2012. Observations on the life history of Enema pan (Fabricius) (Coleoptera: Scarabaeidae: Dynastinae) and its association with bamboo plants (Guadua Kunth: Poaceae: Bambusoideae) in southwestern Amazonia. Coleopterists bulletin, 66: 253–260.
  • Neita, J.C. & Ratcliffe, B.C. 2012. A new species Palaeophileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) from Peru. Coleopterists bulletin, 66: 261–265.
  • Dupuis, F., Neita, J.C. & Ratcliffe, B.C. 2012. Synonymic note on the genus Palaeophileurus Kolbe, 1910 (Coleoptera, Dynastidae). Bulletin de la Société Entomologique de France 117: 460.
  • Ratcliffe, B.C. 2012. Comments on Scarabaeus Linnaeus, 1758, Dynastes MacLeay, 1819, SCARABAEINAE Latreille, 1802, and DYNASTINAE MacLeay, 1819 (Insecta, Coleoptera, SCARABAEOIDEA): proposed conservation of usage (Case 3590) 2. Bulletin of zoological nomenclature, 69: 293–294.















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