Template:Neupane et al., 2015
- Neupane, S., Dessein, S., Wikström, N., Lewis, P.O., Long, C., Bremer, B. & Motley, T.J. 2015. The Hedyotis-Oldenlandia complex (Rubiaceae: Spermacoceae) in Asia and the Pacific: Phylogeny revisited with new generic delimitations. Taxon 64(2): 299-322. DOI: 10.12705/642.8 PDF.