Template:Beutel et al., 2012
- Beutel, R.G., Wang, B., Tan, J.-J., Ge, S.-Q., Ren, D. & Yang, X-K. 2012. On the phylogeny and evolution of Mesozoic and extant lineages of Adephaga (Coleoptera, Insecta). Cladistics 29(2): 147–165. DOI: 10.1111/j.1096-0031.2012.00420.x