Template:Lötters et al., 2009
- Lötters, S., Schmitz, A., Reichle, S., Rödder, D. & Quennet, V. 2009. Another case of cryptic diversity in poison frogs (Dendrobatidae: Ameerega)—description of a new species from Bolivia. Zootaxa 2028: 20–30. Abstract. Full text PDF from ResearchGate.