Template:Helgen et al., 2009

• Helgen, K.M., Cole, F.R., Helgen, L.E. & Wilson, D.E. 2009. Generic Revision in the Holarctic Ground Squirrel Genus Spermophilus. Journal of Mammalogy 90(2): 270–305. DOI: 10.1644/07-MAMM-A-309.1 JSTOR PDF  
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